1.
Groningen (province)
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Groningen is the northeasternmost province of the Netherlands. It borders on Friesland to the west, Drenthe to the south, the German state of Niedersachsen to the east, in 2014, it had a population of 582,640 and a total area of 2,960 km2. The area was part of Frisia, the Frankish Empire, the Holy Roman Empire, and the Dutch Republic. In the 14th century, the city of Groningen became a member of the Hanseatic League, the capital of the province and the seat of the provincial government is the city of Groningen. Since 2016, René Paas has been the Kings Commissioner in the province, a coalition of the Labour Party, Peoples Party for Freedom and Democracy, Democrats 66, and ChristianUnion forms the executive branch. The province is divided into 23 municipalities, the land is mainly used for agriculture. There are sea ports in Delfzijl and Eemshaven, the Groningen gas field was discovered in 1959. The province is home to the University of Groningen and Hanze University of Applied Sciences, Groningen was originally a part of Frisia. It became a part of the Frankish Empire around 785, charlemagne assigned the Christianization of this new possession to Ludger. In the 11th century, the city of Groningen was a village in Drenthe that belonged to the Bishopric of Utrecht, during the Middle Ages, central control was remote, and the city of Groningen acted as a city-state, exerting a dominating influence on the surrounding Ommelanden. In the 14th century, Groningen became one of the towns within the Hanseatic League, in the years after, Groningen expanded its influence. At its peak almost all of the current province Friesland was under the influence, shortly before 1498, Maximilian I, Holy Roman Emperor gave Groningen and Friesland to Albert III, Duke of Saxony, who could however not establish permanent control. In 1514/15 Groningen came to the Duchy of Guelders, and in 1536 as the Lordship of Groningen to the Habsburg Netherlands, in 1594, Groningen was conquered by the Republic of the Seven United Provinces, precursor state of the Netherlands, to which it belonged henceforth. During World War II, the Netherlands were occupied by Nazi Germany, in May 1945, another 3,000 German soldiers were captured in the Battle of Delfzijl by the 5th Canadian Division, after which all of the northern provinces were liberated. East Groningen was the scene of a particularly fierce class struggle in the 19th and 20th centuries, perhaps not coincidentally, Groningen boasts the only municipality where the Communist Party of the Netherlands has ever had a mayor. The northernmost point of the Netherlands is on Rottumerplaat at 53°33′18″N 6°28′41″E, Groningen is the 7th largest province of the Netherlands. It has an area of 2,960 km2, with 2,325 km2 of land and 635 km2 of water. About 80% of the land or 1,876 km2 is used for agriculture
2.
Hybrid (biology)
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In biology, a hybrid, also known as a cross breed, is the result of combining, through sexual reproduction, the qualities of two animals or plants of different breeds, varieties, species or genera. Hybrids are not always intermediate between their parents, but can show heterosis or hybrid vigour, often growing larger or taller than either parent, some act before fertilization, others after it. A few animal species and many plant species, however, are the result of hybrid speciation, doubling the number of chromosomes to create polyploids is important in hybrid speciation, because homoploid hybrids are rarely fertile, the polyploid hybrids are allopolyploids. Human impact on the environment has greatly increased the mixing of species, with introduced species worldwide, the genetic mixing may threaten many species with extinction, while genetic erosion in crop plants may be damaging the gene pools of many species for future breeding. Many commercially useful fruits, flowers, garden herbs and trees have been produced by hybridization, one flower, Oenothera lamarckiana, was central to early genetics research into polyploidy. The term hybrid is derived from Latin hybrida, used for such as of a tame sow and a wild boar, or the child of a freeman. The term came into use in English in the 19th century. Conspicuous hybrids are named with portmanteau words, starting in the 1920s with the breeding of tiger-lion hybrids. The cross between two different homozygous lines produces an F1 hybrid that is heterozygous, having two alleles, one contributed by each parent and typically one is dominant and the other recessive. Typically, the F1 generation is also phenotypically homogeneous, producing offspring that are all similar to each other, double cross hybrids result from the cross between two different F1 hybrids. Three-way cross hybrids result from the cross between an F1 hybrid and an inbred line, triple cross hybrids result from the crossing of two different three-way cross hybrids. Top cross hybrids result from the crossing of a top quality or pure-bred male, population hybrids result from the crossing of plants or animals in a population with those of another population. These include interspecific hybrids or crosses between different breeds. e, from the point of view of genetics, different kinds of hybrid can be distinguished as follows, A genetic hybrid carries two different alleles of the same gene. A structural hybrid results from the fusion of gametes that have differing structure in at least one chromosome, a numerical hybrid results from the fusion of gametes having different haploid numbers of chromosomes. A permanent hybrid results when only the heterozygous genotype occurs, as in Oenothera lamarckiana, from the point of view of taxonomy, hybrids differ according to their parentage, Hybrids between different subspecies are called intra-specific hybrids. Offspring resulting from interspecies mating, are called interspecific hybrids, these result in hybrid speciation. Intergeneric hybrids result from matings between different genera, such as sheep and goats. Interfamilial hybrids such as chickens and guineafowl or pheasants are reliably described but extremely rare
3.
Cultivar
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The term cultivar most commonly refers to an assemblage of plants selected for desirable characteristics that are maintained during propagation. More generally, cultivar refers to the most basic classification category of cultivated plants governed by the ICNCP, most cultivars have arisen in cultivation, but a few are special selections from the wild. Popular ornamental garden plants like roses, camellias, daffodils, rhododendrons, trees used in forestry are also special selections grown for their enhanced quality and yield of timber. Cultivars form a part of Liberty Hyde Baileys broader grouping. Cultivar was coined by Bailey and it is regarded as a portmanteau of cultivated and variety. A cultivar is not the same as a variety, a taxonomic rank below subspecies. In recent times, the naming of cultivars has been complicated by the use of statutory plant patents, the International Union for the Protection of New Varieties of Plants offers legal protection of plant cultivars to people or organisations who introduce new cultivars to commerce. UPOV requires that a cultivar be distinct, uniform and stable, to be distinct, it must have characteristics that easily distinguish it from any other known cultivar. To be uniform and stable, the cultivar must retain these characteristics under repeated propagation, a cultivar is given a cultivar name, which consists of the scientific Latin botanical name followed by a cultivar epithet. The cultivar epithet is usually in a vernacular language, for example, the full cultivar name of the King Edward potato is Solanum tuberosum King Edward. The King Edward part of the name is the cultivar epithet, the origin of the term cultivar arises from the need to distinguish between wild plants and those with characteristics that have arisen in cultivation. This distinction dates back to the Greek philosopher Theophrastus, the Father of Botany, botanical historian Alan Morton notes that Theophrastus in his Enquiry into Plants had an inkling of the limits of culturally induced changes and of the importance of genetic constitution. In Species Plantarum, Linnaeus listed all the known to him. Most of the listed by Linnaeus were of garden origin rather than being wild plants. Over time there was an increasing need to distinguish between plants growing in the wild, and those with variations that had produced in cultivation. In the nineteenth century many garden-derived plants were given names, sometimes in Latin. In the twentieth century an improved international terminology was proposed for the classification and it is essentially the equivalent of the botanical variety except in respect to its origin. However, Bailey was never explicit about the etymology of the word, and it has suggested that it is a contraction of the words cultigen and variety
4.
Ulmus glabra
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A large, deciduous tree, it is essentially a montane species, growing at elevations of up to 1500 m, preferring sites with moist soils and high humidity. The tree can form pure forests in Scandinavia and occurs as far north as latitude 67°N at Beiarn in Norway, wych elm has also been successfully introduced to Narsarsuaq, near the southern tip of Greenland. The tree was by far the most common elm in the north, however, this is contested, the correct Gaelic name considered Loch Laomainn, its origin obscure. The word wych comes from the Old English wice, meaning pliant or supple, an older name for the tree was wych hazel, perhaps based on the similar appearance of hazel and wych elm shoots. Some botanists, notably Lindquist have proposed dividing the species into two subspecies, Ulmus glabra subsp, glabra in the south of the species range, with broad leaves, trees often with a short, forked trunk and a low, broad crown Ulmus glabra subsp. Montana Lindqvist in the north of the range, the leaves are narrower, trees commonly with a long single trunk. The wych elm sometimes reaches heights of 40 m, typically with a crown where open-grown. There are not normally root suckers, natural reproduction is by seed alone, the tree is notable for its very tough, supple young shoots, which are always without the corky ridges or wings characteristic of many elms. The alternate leaves are deciduous, 6–17 cm long by 3–12 cm broad, usually obovate with a base, the lobe often completely covering the short petiole. Leaves on juvenile or shade-grown shoots sometimes have three or more lobes near the apex, the perfect hermaphrodite flowers appear before the leaves in early spring, produced in clusters of 10–20, they are 4 mm across on 10 mm long stems and, being wind-pollinated, are apetalous. The fruit is a winged samara 20 mm long and 15 mm broad, with a single, round,6 mm seed in the centre, maturing in late spring. Montana While the species is susceptible to Dutch elm disease. Research in Spain has indicated the presence of a triterpene, alnulin, moreover, once the tree is dying, its bark is quickly colonized by the fungus Phoma, which radically reduces the amount of bark available for the beetle to breed on. In European trials, clones of apparently resistant trees were inoculated with the pathogen, DNA analysis by Cemagref in France has determined the genetic diversity within the species is very limited, making the chances of a resistant tree evolving rather remote. The Swedish Forest Tree Breeding Association at Källstorp produced triploid and tetraploid forms of the tree, in trials conducted in Italy, the tree was found to have a slight to moderate susceptibility to elm yellows, and a high susceptibility to the elm leaf beetle Xanthogaleruca luteola. The wych elm is moderately shade-tolerant, but requires deep, rich soils as typically found along river valleys, however, the species is intolerant of flooding, as it is of prolonged drought. Although rarely used as a street tree owing to its shape, it can be tolerant of urban air pollution, constricted growing conditions. As wych elm does not sucker from the roots, and any seedlings are often consumed by uncontrolled deer populations, the resultant decline has been extreme, and the wych elm is now uncommon over much of its former range
5.
Ulmus minor
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Ulmus minor Mill. the field elm, is by far the most polymorphic of the European species, although its taxonomy remains a matter of contention. The trees typical habitat is low-lying forest along the rivers, growing in association with oak and ash. Current treatment of the species owes much to Richens, who noted that several varieties of elm are distinguishable on the European mainland. As for British varieties, the continental populations most closely related are in central Europe and he concluded, however, that owing to incomplete field-research at the time of writing, it was not possible to present an overall breakdown of the European Field Elm into regional varieties. Richens sank a number of British elms, notably English elm, however, Melville, writing ten years later, identified five distinct species, several varieties and numerous complex hybrids. In 1992 Armstrong identified no fewer than forty British species and microspecies, clive Stace wrote of the British elms The two-species concept of Richens is not sufficiently discriminating to be of taxonomic value. Nevertheless, it is Richens’ classification which has been the most commonly adopted in recent years, in 2009 Dr Max Coleman of the Royal Botanic Garden, Edinburgh wrote, The advent of DNA fingerprinting has shed considerable light on the question. This means that enigmatic British elms such as Plot elm and English elm have been shown to be clones of field elm. Although Richens did not have the evidence to prove it, he was correct in recognising a series of clones and it is hoped that analysis of molecular markers will ultimately eliminate the taxonomic confusion. The tree typically grows to <30 m and bears a rounded crown, the bark of the trunk is rough, furrowed lightly in older trees to form a block pattern. Young branchlets occasionally have corky wings, the shoots are slender compared with those of wych elm. The leaves are smaller than those of the other European species, hence the specific epithet minor, a common characteristic is the presence of minute black glands along the leaf veins, detectable with the aid of a magnifying glass. The samarae are ovate and notched, the notch extending to the central seed. The species readily produces suckers from roots and stumps, even after devastation by Dutch elm disease, most trees are very susceptible to Dutch elm disease, including all the fashionable pre-20th century plantsmans clones. In the UK, despite its late leaf-flush in the north and its suckering habits, augustine Henry wrote in 1913 that the U. minor planted in parks in Scotland were of French origin. More recently U. minor seed was imported to the UK from Italy, there are mature survivors in Edinburgh that are not the common U. minor cultivars. U. minor has been introduced to the hemisphere, notably Australasia. U. minor can live to a great age, an ancient field elm stood until recently in the village square of Metaxades, Thrace, Greece
6.
Glossary of leaf morphology
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The following is a defined list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple or compound, the edge of the leaf may be regular or irregular, may be smooth or bearing hair, bristles or spines. For more terms describing other aspects of leaves besides their overall morphology see the leaf article, leaves of most plants include a flat structure called the blade or lamina, but not all leaves are flat, some are cylindrical. Leaves may be simple, with a leaf blade, or compound. In flowering plants, as well as the blade of the leaf, there may be a petiole and stipules, leaf structure is described by several terms that include, Being one of the more visible features, leaf shape is commonly used for plant identification. Edge and margin are both interchangeable in the sense that they refer to the perimeter of a leaf. Leaves may also be folded or rolled in various ways, the folding of leaves within a bud is vernation, ptyxis is the folding of an individual leaf in a bud
7.
Dutch elm disease
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Dutch elm disease is caused by a member of the sac fungi affecting elm trees, and is spread by elm bark beetles. It has also reached New Zealand, the disease affects species in the genera Ulmus and Zelkova, therefore it is not specific to the Dutch elm hybrid. The causative agents of DED are ascomycete microfungi, three species are now recognized, Ophiostoma ulmi, which afflicted Europe from 1910, reaching North America on imported timber in 1928. Ophiostoma himal-ulmi, an endemic to the western Himalaya. Ophiostoma novo-ulmi, an extremely virulent species from Japan which was first described in Europe, DED is spread in North America by three species of bark beetles, The native elm bark beetle, Hylurgopinus rufipes. The European elm bark beetle, Scolytus multistriatus, the banded elm bark beetle, Scolytus schevyrewi. In Europe, while S. multistriatus still acts as a vector for infection, it is less effective than the large elm bark beetle. H. rufipes can be a vector for the disease, but is inefficient compared to the other vectors, S. schevyrewi was found in 2003 in Colorado and Utah. Other reported DED vectors include Scolytus sulcifrons, S. pygmaeus, S. laevis, Pteleobius vittatus, other elm bark beetle species are also likely vectors. In an attempt to block the fungus from spreading farther, the tree reacts by plugging its own xylem tissue with gum and tyloses, bladder-like extensions of the xylem cell wall. As the xylem delivers water and nutrients to the rest of the plant, the first sign of infection is usually an upper branch of the tree with leaves starting to wither and yellow in summer, months before the normal autumnal leaf shedding. This progressively spreads to the rest of the tree, with further dieback of branches, eventually, the roots die, starved of nutrients from the leaves. Often, not all the roots die, the roots of some species, notably the English elm Ulmus procera, put up suckers which flourish for approximately 15 years, after which they too succumb. Dutch elm disease was first noticed in continental Europe in 1910, the disease was isolated in The Netherlands in 1921 by Bea Schwarz, a pioneering Dutch phytopathologist, and this discovery would lend the disease its name. Circa 1967, a new, far more virulent strain arrived in Britain on a shipment of rock elm U, the disease is still migrating northwards through Scotland, reaching Edinburgh in the late 1970s, and Inverness in 2006. By 1990, very few mature elms were left in Britain or much of continental Europe, one of the most distinctive English countryside trees, the English elm U. procera Salisb. is particularly susceptible. Thirty years after the outbreak of the epidemic, nearly all these trees, the species still survives in hedgerows, as the roots are not killed and send up root sprouts. These suckers rarely reach more than 5 m tall before succumbing to a new attack of the fungus, however, established hedges kept low by clipping have remained apparently healthy throughout the nearly 40 years since the onset of the disease in the UK
8.
William Dampier
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William Dampier was an English explorer and navigator who became the first Englishman to explore parts of what is today Australia, and the first person to circumnavigate the world three times. He has also described as Australias first natural historian, as well as one of the most important British explorers of the period between Sir Walter Raleigh and James Cook. On a later voyage he rescued Alexander Selkirk, a former crewmate who may have inspired Daniel Defoes Robinson Crusoe, others influenced by Dampier include James Cook, Horatio Nelson, Charles Darwin, and Alfred Russel Wallace. William Dampier was born at Hymerford House in East Coker, Somerset and he was baptised on 5 September, but his precise date of birth is not recorded. He was educated at Kings School, Bruton, Dampier sailed on two merchant voyages to Newfoundland and Java before joining the Royal Navy in 1673. He took part in the two Battles of Schooneveld in June of that year, Dampiers service was cut short by a catastrophic illness, and he returned to England for several months of recuperation. For the next years he tried his hand at various careers, including plantation management in Jamaica and logging in Mexico. Returning to England, he married Judith around 1679, only to leave for the sea a few months later. This led to his first circumnavigation, during which he accompanied a raid across the Isthmus of Darién in Panama, the pirates then raided Spanish settlements in Peru before returning to the Caribbean. Dampier made his way to Virginia, where in 1683 he was engaged by the privateer John Cooke, Cooke entered the Pacific via Cape Horn and spent a year raiding Spanish possessions in Peru, the Galápagos Islands, and Mexico. This expedition collected buccaneers and ships as it went along, at one time having a fleet of ten vessels, Cooke died in Mexico, and a new leader, Edward Davis, was elected captain by the crew. Dampier transferred to the privateer Charles Swans ship, Cygnet, and on 31 March 1686 they set out across the Pacific to raid the East Indies, calling at Guam, Spanish witnesses saw the predominantly English crew as not only pirates and heretics but also cannibals. Leaving Swan and 36 others behind on Mindanao, the rest of the privateers sailed on to Manila, Poulo Condor, China, the Spice Islands, and New Holland. On 5 January 1688, Cygnet anchored two miles from shore in 29 fathoms on the northwest coast of Australia, near King Sound. Dampier and his ship remained there until March 12, and while the ship was being careened Dampier made notes on the fauna and flora, among his fellows were a significant number of Spanish sailors, most notably Alonso Ramírez, a native of San Juan, Puerto Rico. Later that year, by agreement, Dampier and two shipmates were marooned on one of the Nicobar Islands and they obtained a small canoe which they modified after first capsizing and then, after surviving a great storm at sea, called at Acheen in Sumatra. Dampier returned to England in 1691 via the Cape of Good Hope, penniless and he also had as a source of income a slave known as Prince Jeoly, from Miangas, who became famous for his tattoos. Dampier exhibited Jeoly in London, thereby also generating publicity for a book based on his diaries, the publication of the book, A New Voyage Round the World, in 1697 was a popular sensation, creating interest at the Admiralty
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