Verrucariaceae is a family of lichens and a few non-lichenised fungi in the order Verrucariales. The lichens have a wide variety of thallus forms, from crustose (crust-like) to foliose (bushy) and squamulose (scaly). Most of them grow on land, some in freshwater and a few in the sea. Many are free-living but there are some species that are parasites on other lichens, while one marine species always lives together with a leafy green alga.
Verrucariaceae
Dermatocarpon americanum is a foliose member of the Verrucariaceae ...
... while Staurothele pulvinata is squamulose.
Catapyrenium boccanum, shown here growing in a hole in a calcareous rock, is a catapyrenioid lichen.
Lichens are symbiotic organisms made up of multiple species: a fungus, one or more photobionts and sometimes a yeast. They are regularly grouped by their external appearance – a characteristic known as their growth form. This form, which is based on the appearance of vegetative part of the lichen, varies depending on the species and the environmental conditions it faces. Those who study lichens (lichenologists) have described a dozen of these forms: areolate, byssoid, calicioid, cladoniform, crustose, filamentous, foliose, fruticose, gelatinous, leprose, placoidioid and squamulose. Traditionally, crustose (flat), foliose (leafy) and fruticose (shrubby) are considered to be the three main forms. In addition to these more formalised, traditional growth types, there are a handful of informal types named for their resemblance to the lichens of specific genera. These include alectorioid, catapyrenioid, cetrarioid, hypogymnioid, parmelioid and usneoid.
Crustose (Lecidella elaeochroma)
Foliose (Flavoparmelia caperata)
Fruticose (Usnea filipendula)
Rhizocarpon species