1.
Taxonomy (biology)
–
Taxonomy is the science of defining groups of biological organisms on the basis of shared characteristics and giving names to those groups. The exact definition of taxonomy varies from source to source, but the core of the remains, the conception, naming. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy, the broadest meaning of taxonomy is used here. The word taxonomy was introduced in 1813 by Candolle, in his Théorie élémentaire de la botanique, the term alpha taxonomy is primarily used today to refer to the discipline of finding, describing, and naming taxa, particularly species. In earlier literature, the term had a different meaning, referring to morphological taxonomy, ideals can, it may be said, never be completely realized. They have, however, a value of acting as permanent stimulants. Some of us please ourselves by thinking we are now groping in a beta taxonomy, turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy, thus, Ernst Mayr in 1968 defined beta taxonomy as the classification of ranks higher than species. This activity is what the term denotes, it is also referred to as beta taxonomy. How species should be defined in a group of organisms gives rise to practical and theoretical problems that are referred to as the species problem. The scientific work of deciding how to define species has been called microtaxonomy, by extension, macrotaxonomy is the study of groups at higher taxonomic ranks, from subgenus and above only, than species. While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, earlier works were primarily descriptive, and focused on plants that were useful in agriculture or medicine. There are a number of stages in scientific thinking. Early taxonomy was based on criteria, the so-called artificial systems. Later came systems based on a complete consideration of the characteristics of taxa, referred to as natural systems, such as those of de Jussieu, de Candolle and Bentham. The publication of Charles Darwins Origin of Species led to new ways of thinking about classification based on evolutionary relationships and this was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler and Engler, the advent of molecular genetics and statistical methodology allowed the creation of the modern era of phylogenetic systems based on cladistics, rather than morphology alone. Taxonomy has been called the worlds oldest profession, and naming and classifying our surroundings has likely been taking place as long as mankind has been able to communicate
Taxonomy (biology)
–
Title page of
Systema Naturae, Leiden, 1735
Taxonomy (biology)
–
Evolution of the
vertebrates at class level, width of spindles indicating number of families. Spindle diagrams are typical for
Evolutionary taxonomy
Taxonomy (biology)
–
The same relationship, expressed as a
cladogram typical for
cladistics
Taxonomy (biology)
–
Type specimen for
Nepenthes smilesii, a tropical
pitcher plant.
2.
Plantae
–
Plants are mainly multicellular, predominantly photosynthetic eukaryotes of the kingdom Plantae. The term is generally limited to the green plants, which form an unranked clade Viridiplantae. This includes the plants, conifers and other gymnosperms, ferns, clubmosses, hornworts, liverworts, mosses and the green algae. Green plants have cell walls containing cellulose and obtain most of their energy from sunlight via photosynthesis by primary chloroplasts and their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic and have lost the ability to produce amounts of chlorophyll or to photosynthesize. Plants are characterized by sexual reproduction and alternation of generations, although reproduction is also common. There are about 300–315 thousand species of plants, of which the great majority, green plants provide most of the worlds molecular oxygen and are the basis of most of Earths ecologies, especially on land. Plants that produce grains, fruits and vegetables form humankinds basic foodstuffs, Plants play many roles in culture. They are used as ornaments and, until recently and in variety, they have served as the source of most medicines. The scientific study of plants is known as botany, a branch of biology, Plants are one of the two groups into which all living things were traditionally divided, the other is animals. The division goes back at least as far as Aristotle, who distinguished between plants, which generally do not move, and animals, which often are mobile to catch their food. Much later, when Linnaeus created the basis of the system of scientific classification. Since then, it has become clear that the plant kingdom as originally defined included several unrelated groups, however, these organisms are still often considered plants, particularly in popular contexts. When the name Plantae or plant is applied to a group of organisms or taxon. The evolutionary history of plants is not yet settled. Those which have been called plants are in bold, the way in which the groups of green algae are combined and named varies considerably between authors. Algae comprise several different groups of organisms which produce energy through photosynthesis, most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble land plants, but are classified among the brown, red and green algae. Each of these groups also includes various microscopic and single-celled organisms
Plantae
–
Green algae from
Ernst Haeckel 's
Kunstformen der Natur, 1904.
Plantae
–
Had'n
Plantae
–
Dicksonia antarctica, a species of
tree fern
Plantae
–
A petrified log in
Petrified Forest National Park.
3.
Angiosperms
–
The flowering plants, also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 416 families, approx. 13,164 known genera and a total of c.295,383 known species, etymologically, angiosperm means a plant that produces seeds within an enclosure, in other words, a fruiting plant. The term angiosperm comes from the Greek composite word meaning enclosed seeds, the ancestors of flowering plants diverged from gymnosperms in the Triassic Period, during the range 245 to 202 million years ago, and the first flowering plants are known from 160 mya. They diversified extensively during the Lower Cretaceous, became widespread by 120 mya, angiosperms differ from other seed plants in several ways, described in the table. These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants, the amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms. The vascular bundles of the stem are arranged such that the xylem and phloem form concentric rings, in the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium, the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Among the monocotyledons, the bundles are more numerous in the stem and are scattered through the ground tissue. They contain no cambium and once formed the stem increases in diameter only in exceptional cases, the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, and provide the most trustworthy external characteristics for establishing relationships among angiosperm species, the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally on a shoot or from the axil of a leaf, occasionally, as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. There are two kinds of cells produced by flowers. Microspores, which divide to become pollen grains, are the male cells and are borne in the stamens. The female cells called megaspores, which divide to become the egg cell, are contained in the ovule. The flower may consist only of parts, as in willow. Usually, other structures are present and serve to protect the sporophylls, the individual members of these surrounding structures are known as sepals and petals. The outer series is usually green and leaf-like, and functions to protect the rest of the flower, the inner series is, in general, white or brightly colored, and is more delicate in structure. It functions to attract insect or bird pollinators, attraction is effected by color, scent, and nectar, which may be secreted in some part of the flower
Angiosperms
Angiosperms
–
Bud of a pink
rose
Angiosperms
–
Flowers of
Malus sylvestris (crab apple)
Angiosperms
–
Flowers and leaves of
Oxalis pes-caprae (Bermuda buttercup)
4.
Monocots
–
Monocotyledons, commonly referred to as monocots, are flowering plants whose seeds typically contain only one embryonic leaf, or cotyledon. However, molecular research has shown that while the monocots form a monophyletic group or clade. Monocots have almost always recognized as a group, but with various taxonomic ranks. The APG III system of 2009 recognises a clade called monocots, the monocots include about 60,000 species. The largest family in this group by number of species are the orchids, about half as many species belong to the true grasses, which are economically the most important family of monocots. In agriculture the majority of the biomass produced comes from monocots and these include not only major grains, but also forage grasses, sugar cane, and the bamboos. Other economically important monocot crops include various palms, bananas, gingers and their relatives, turmeric and cardamom, asparagus and the onions and garlic family. Additionally most of the bulbs, plants cultivated for their blooms, such as lilies, daffodils, irises, amaryllis, cannas, bluebells. The monocots or monocotyledons have, as the name implies, a single cotyledon, or embryonic leaf, from a diagnostic point of view the number of cotyledons is neither a particularly useful characteristic, nor is it completely reliable. Nevertheless, monocots are sufficiently distinctive that there has rarely been disagreement as to membership of this group, however, morphological features that reliably characterise major clades are rare. Thus monocots are distinguishable from other angiosperms both in terms of their uniformity and diversity, although largely herbaceous, some arboraceous monocots reach great height, length and mass. The latter include agaves, palms, pandans, and bamboos and this creates challenges in water transport that monocots deal with in various ways. Some such as species of Yucca develop anomalous secondary growth, while palm trees, the axis undergoes primary thickening, that progresses from internode to internode, resulting in a typical inverted conical shape of the basal primary axis. The limited conductivity also contributes to limited branching of the stems, despite these limitations a wide variety of adaptive growth forms has resulted from epiphytic orchids and bromeliads to submarine Alismatales and mycotrophic Burmanniaceae and Triuridaceae. Other monocots, particularly Poales have adopted a life form. Leaves The cotyledon, the primordial Angiosperm leaf consists of a proximal leaf base or hypophyll, in moncots the hypophyll tends to be the dominant part in contrast to other angiosperms. Mature monocot leaves are narrow and linear, forming a sheathing around the stem at its base. There is usually only one leaf per node because the leaf base encompasses more than half the circumference, the evolution of this monocot characteristic has been attributed to developmental differences in early zonal differentiation rather than meristem activity
Monocots
Monocots
–
Comparison of a monocot and dicot sprouting. Note that the visible part of the monocot plant (left) is actually the first true leaf produced from the
meristem; the cotyledon itself remains within the seed.
Monocots
–
Allium crenulatum, an onion, with a typical monocot
perianth and parallel leaf venation
Monocots
–
Slice of onion, showing parallel veins in cross section
5.
Commelinids
–
In plant taxonomy, commelinids is a name used by the APG III system for a clade within the monocots, which in its turn is a clade within the angiosperms. The commelinids are the clade that the APG has informally named within the monocots. The remaining monocots are a paraphyletic unit, also known as the commelinid monocots it forms one of three groupings within the monocots, and the final branch, the other two groups being the alismatid monocots and the lilioid monocots. Members of the clade have cell walls containing UV-fluorescent ferulic acid. The commelinids were first recognized as a group in 1967 by Armen Takhtajan. However, by the release of his 1980 system of classification, the commelinids constitute a well-supported clade within the monocots, and this clade has been recognized in all four APG classification systems. The commelinids of APG II and APG III contain essentially the same plants as the commelinoids of the earlier APG system, in APG IV the family Dasypogonaceae is no longer directly placed under commelinids but instead a family of order Arecales. Media related to Commelinids at Wikimedia Commons
Commelinids
–
commelinids
6.
Poales
–
The Poales are a large order of flowering plants in the monocotyledons, and includes families of plants such as the grasses, bromeliads, and sedges. Sixteen plant families are recognized by botanists to be part of Poales. The flowers are small, enclosed by bracts, and arranged in inflorescences. The flowers of species are wind pollinated, the seeds usually contain starch. It did not include the Bromeliaceae and Mayaceae, but had the additional families Prioniaceae, Sparganiaceae, the morphology-based Cronquist system did not include an order named Poales, assigning these families to the orders Bromeliales, Cyperales, Hydatellales, Juncales, Restionales and Typhales. The earliest known fossils include pollen and fruits, major lineages within the Poales have been referred to as bromeliad, cyperid, xyrid, graminid, and restiid clades. A phylogenetic analysis resolved most relationships within the order but found support for the monophyly of the cyperid clade. The relationship between Centrolepidaceae and Restoniaceae within the clade remains unclear, the first may actually be embedded in the latter. Within the order, by far the most important family economically is the family of grasses, which includes barley, maize, millet, rice, many Bromeliaceae are used as ornamental plants. Bremer, K. Gondwanan Evolution of the Grass Alliance of Families, judd, W. S. C. S. Campbell, E. A. Kellogg, P. F. Stevens, M. J. Donoghue. Plant Systematics, A Phylogenetic Approach, 2nd edition, linder, H. Peter, Rudall, Paula J. Evolutionary History of the Poales. Annual Review of Ecology, Evolution, and Systematics, flora of the Southeastern United States,48
Poales
Poales
–
Setaria verticillata from family
Poaceae
Poales
–
Billbergia pyramidalis of family
Bromeliaceae
7.
Poaceae
–
Poaceae or Gramineae is a large and nearly ubiquitous family of monocotyledonous flowering plants known as grasses. Poaceae includes the cereal grasses, bamboos and the grasses of natural grassland and cultivated lawns, Grasses have stems that are hollow except at the nodes and narrow alternate leaves borne in two ranks. The lower part of each leaf encloses the stem, forming a leaf-sheath, with ca 780 genera and around 12,000 species, Poaceae are the fifth-largest plant family, following the Asteraceae, Orchidaceae, Fabaceae and Rubiaceae. Grasslands such as savannah and prairie grasses are dominant are estimated to constitute 40. 5% of the land area of the Earth, excluding Greenland. Grasses are also an important part of the vegetation in many habitats, including wetlands, forests. Though commonly called grasses, seagrasses, rushes, and sedges fall outside this family, the rushes and sedges are related to the Poaceae, being members of the order Poales, but the seagrasses are members of order Alismatales. The name Poaceae was given by John Hendley Barnhart in 1895, based on the tribe Poeae described in 1814 by Robert Brown, the term is derived from the Ancient Greek πόα. Grasses include some of the most versatile plant life-forms, a cladogram shows subfamilies and approximate species numbers in brackets, Before 2005, fossil findings indicated that grasses evolved around 55 million years ago. Recent findings of grass-like phytoliths in Cretaceous dinosaur coprolites have pushed this back to 66 million years ago. In 2011, revised dating of the origins of the rice tribe Oryzeae suggested a date as early as 107 to 129 Mya, a multituberculate mammal with grass-eating adaptations seems to suggest that grasses were already around at 120 mya. This separation occurred within the short time span of about 4 million years. Grass leaves are always alternate and distichous, and have parallel veins. Each leaf is differentiated into a lower sheath hugging the stem, the leaf blades of many grasses are hardened with silica phytoliths, which discourage grazing animals, some, such as sword grass, are sharp enough to cut human skin. A membranous appendage or fringe of hairs called the ligule lies at the junction between sheath and blade, preventing water or insects from penetrating into the sheath, flowers of Poaceae are characteristically arranged in spikelets, each having one or more florets. The spikelets are further grouped into panicles or spikes, the part of the spikelet that bears the florets is called the rachilla. A spikelet consists of two bracts at the base, called glumes, followed by one or more florets, a floret consists of the flower surrounded by two bracts, one external—the lemma—and one internal—the palea. The flowers are usually hermaphroditic—maize being an important exception—and anemophilous or wind-pollinated, the perianth is reduced to two scales, called lodicules, that expand and contract to spread the lemma and palea, these are generally interpreted to be modified sepals. This complex structure can be seen in the image on the right, the fruit of grasses is a caryopsis, in which the seed coat is fused to the fruit wall
Poaceae
Poaceae
–
Grass flowers
Poaceae
–
Setaria verticillata from
Panicoideae
Poaceae
–
Tragus roxburghii from
Chloridoideae
8.
Triticum
–
Wheat is a cereal grain, originally from the Levant region but now cultivated worldwide. In 2016, world production of wheat was 749 million tonnes, making it the second most-produced cereal after maize, since 1960, world production of wheat and other grain crops has tripled and is expected to grow further through the middle of the 21st Century. This grain is grown on land area than any other commercial food. World trade in wheat is greater than for all other crops combined, globally, wheat is the leading source of vegetal protein in human food, having a protein content of about 13%, which is relatively high compared to other major cereals and staple foods. The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BCE. In a small part of the population, gluten – the major part of wheat protein – can trigger coeliac disease, non-coeliac gluten sensitivity, gluten ataxia. Cultivation and repeated harvesting and sowing of the grains of wild grasses led to the creation of domestic strains, in domesticated wheat, grains are larger, and the seeds remain attached to the ear by a toughened rachis during harvesting. In wild strains, a more fragile rachis allows the ear to easily shatter, as the traits that improve wheat as a food source also involve the loss of the plants natural seed dispersal mechanisms, highly domesticated strains of wheat cannot survive in the wild. Cultivation of wheat began to spread beyond the Fertile Crescent after about 8000 BCE, jared Diamond traces the spread of cultivated emmer wheat starting in the Fertile Crescent sometime before 8800 BCE. Archaeological analysis of wild emmer indicates that it was first cultivated in the southern Levant with finds dating back as far as 9600 BCE, Genetic analysis of wild einkorn wheat suggests that it was first grown in the Karacadag Mountains in southeastern Turkey. Dated archeological remains of wheat in settlement sites near this region, including those at Abu Hureyra in Syria. With the anomalous exception of two grains from Iraq ed-Dubb, the earliest carbon-14 date for einkorn wheat remains at Abu Hureyra is 7800 to 7500 years BCE. Remains of harvested emmer from several sites near the Karacadag Range have been dated to between 8600 and 8400 BCE, that is, in the Neolithic period and these remains were dated by Willem van Zeist and his assistant Johanna Bakker-Heeres to 8800 BCE. They also concluded that the settlers of Tell Aswad did not develop this form of emmer themselves, the cultivation of emmer reached Greece, Cyprus and India by 6500 BCE, Egypt shortly after 6000 BCE, and Germany and Spain by 5000 BCE. The early Egyptians were developers of bread and the use of the oven, by 3000 BCE, wheat had reached the British Isles and Scandinavia. A millennium later it reached China, the oldest evidence for hexaploid wheat has been confirmed through DNA analysis of wheat seeds, dating to around 6400-6200 BCE, recovered from Çatalhöyük. The first identifiable bread wheat with sufficient gluten for yeasted breads has been identified using DNA analysis in samples from a dating to approximately 1350 BCE at Assiros in Macedonia. From Asia, wheat continued to spread throughout Europe, in the British Isles, wheat straw was used for roofing in the Bronze Age, and was in common use until the late 19th century
Triticum
–
Wheat
Triticum
–
Spikelets of a hulled wheat,
einkorn
Triticum
–
Green wheat a month before harvest
Triticum
–
Wheat harvest on the
Palouse,
Idaho, United States
9.
Binomial nomenclature
–
Such a name is called a binomial name, a binomen, binominal name or a scientific name, more informally it is also called a Latin name. The first part of the name identifies the genus to which the species belongs, for example, humans belong to the genus Homo and within this genus to the species Homo sapiens. The formal introduction of system of naming species is credited to Carl Linnaeus. But Gaspard Bauhin, in as early as 1623, had introduced in his book Pinax theatri botanici many names of genera that were adopted by Linnaeus. Although the general principles underlying binomial nomenclature are common to these two codes, there are differences, both in the terminology they use and in their precise rules. Similarly, both parts are italicized when a binomial name occurs in normal text, thus the binomial name of the annual phlox is now written as Phlox drummondii. In scientific works, the authority for a name is usually given, at least when it is first mentioned. In zoology Patella vulgata Linnaeus,1758, the original name given by Linnaeus was Fringilla domestica, the parentheses indicate that the species is now considered to belong in a different genus. The ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, in botany Amaranthus retroflexus L. – L. is the standard abbreviation used in botany for Linnaeus. – Linnaeus first named this bluebell species Scilla italica, Rothmaler transferred it to the genus Hyacinthoides, the ICN does not require that the dates of either publication be specified. Prior to the adoption of the binomial system of naming species. Together they formed a system of polynomial nomenclature and these names had two separate functions. First, to designate or label the species, and second, to be a diagnosis or description, such polynomial names may sometimes look like binomials, but are significantly different. For example, Gerards herbal describes various kinds of spiderwort, The first is called Phalangium ramosum, Branched Spiderwort, is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia. The Latin phrases are short descriptions, rather than identifying labels, the Bauhins, in particular Caspar Bauhin, took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of binomial nomenclature is due to Swedish botanist and physician Carl von Linné. It was in his 1753 Species Plantarum that he first began using a one-word trivial name together with a generic name in a system of binomial nomenclature. This trivial name is what is now known as an epithet or specific name
Binomial nomenclature
–
Carl Linnaeus (1707–1778), a Swedish botanist, invented the modern system of binomial nomenclature.
10.
Carl Linnaeus
–
Carl Linnaeus, also known after his ennoblement as Carl von Linné, was a Swedish botanist, physician, and zoologist, who formalised the modern system of naming organisms called binomial nomenclature. He is known by the father of modern taxonomy. Many of his writings were in Latin, and his name is rendered in Latin as Carolus Linnæus, Linnaeus was born in the countryside of Småland, in southern Sweden. He received most of his education at Uppsala University. He lived abroad between 1735 and 1738, where he studied and also published a first edition of his Systema Naturae in the Netherlands and he then returned to Sweden, where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on journeys through Sweden to find and classify plants. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, at the time of his death, he was one of the most acclaimed scientists in Europe. The philosopher Jean-Jacques Rousseau sent him the message, Tell him I know no man on earth. The German writer Johann Wolfgang von Goethe wrote, With the exception of Shakespeare and Spinoza, Swedish author August Strindberg wrote, Linnaeus was in reality a poet who happened to become a naturalist. Among other compliments, Linnaeus has been called Princeps botanicorum, The Pliny of the North and he is also considered as one of the founders of modern ecology. In botany, the abbreviation used to indicate Linnaeus as the authority for species names is L. In older publications, sometimes the abbreviation Linn. is found, Linnæus was born in the village of Råshult in Småland, Sweden, on 23 May 1707. He was the first child of Nicolaus Ingemarsson and Christina Brodersonia and his siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus, and Emerentia Linnæa. One of a line of peasants and priests, Nils was an amateur botanist, a Lutheran minister. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius, a year after Linnæus birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curates house, even in his early years, Linnæus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus, soon Linnæus was given his own patch of earth where he could grow plants
Carl Linnaeus
–
Carl von Linné,
Alexander Roslin, 1775. Oil painting in the portrait collection at
Gripsholm Castle
Carl Linnaeus
–
The
coat of arms of Carl von Linné.
Carl Linnaeus
–
Birthplace of Linnæus at
Råshult
Carl Linnaeus
–
Statue of Linnaeus as a University student in
Lund
11.
Wheat
–
Wheat is a cereal grain, originally from the Levant region but now cultivated worldwide. In 2016, world production of wheat was 749 million tonnes, making it the second most-produced cereal after maize, since 1960, world production of wheat and other grain crops has tripled and is expected to grow further through the middle of the 21st Century. This grain is grown on land area than any other commercial food. World trade in wheat is greater than for all other crops combined, globally, wheat is the leading source of vegetal protein in human food, having a protein content of about 13%, which is relatively high compared to other major cereals and staple foods. The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BCE. In a small part of the population, gluten – the major part of wheat protein – can trigger coeliac disease, non-coeliac gluten sensitivity, gluten ataxia. Cultivation and repeated harvesting and sowing of the grains of wild grasses led to the creation of domestic strains, in domesticated wheat, grains are larger, and the seeds remain attached to the ear by a toughened rachis during harvesting. In wild strains, a more fragile rachis allows the ear to easily shatter, as the traits that improve wheat as a food source also involve the loss of the plants natural seed dispersal mechanisms, highly domesticated strains of wheat cannot survive in the wild. Cultivation of wheat began to spread beyond the Fertile Crescent after about 8000 BCE, jared Diamond traces the spread of cultivated emmer wheat starting in the Fertile Crescent sometime before 8800 BCE. Archaeological analysis of wild emmer indicates that it was first cultivated in the southern Levant with finds dating back as far as 9600 BCE, Genetic analysis of wild einkorn wheat suggests that it was first grown in the Karacadag Mountains in southeastern Turkey. Dated archeological remains of wheat in settlement sites near this region, including those at Abu Hureyra in Syria. With the anomalous exception of two grains from Iraq ed-Dubb, the earliest carbon-14 date for einkorn wheat remains at Abu Hureyra is 7800 to 7500 years BCE. Remains of harvested emmer from several sites near the Karacadag Range have been dated to between 8600 and 8400 BCE, that is, in the Neolithic period and these remains were dated by Willem van Zeist and his assistant Johanna Bakker-Heeres to 8800 BCE. They also concluded that the settlers of Tell Aswad did not develop this form of emmer themselves, the cultivation of emmer reached Greece, Cyprus and India by 6500 BCE, Egypt shortly after 6000 BCE, and Germany and Spain by 5000 BCE. The early Egyptians were developers of bread and the use of the oven, by 3000 BCE, wheat had reached the British Isles and Scandinavia. A millennium later it reached China, the oldest evidence for hexaploid wheat has been confirmed through DNA analysis of wheat seeds, dating to around 6400-6200 BCE, recovered from Çatalhöyük. The first identifiable bread wheat with sufficient gluten for yeasted breads has been identified using DNA analysis in samples from a dating to approximately 1350 BCE at Assiros in Macedonia. From Asia, wheat continued to spread throughout Europe, in the British Isles, wheat straw was used for roofing in the Bronze Age, and was in common use until the late 19th century
Wheat
–
Wheat
Wheat
–
Spikelets of a hulled wheat,
einkorn
Wheat
–
Green wheat a month before harvest
Wheat
–
Wheat harvest on the
Palouse,
Idaho, United States
12.
Diploid
–
Ploidy is the number of sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Cells are described according to the number of present, monoploid, diploid, triploid, tetraploid, pentaploid, hexaploid, heptaploid or septaploid. The generic term polyploid is used to describe cells with three or more sets of chromosomes, humans are diploid organisms, carrying two complete sets of chromosomes, one set of 23 chromosomes from their father and one set of 23 chromosomes from their mother. The two sets combined provide a complement of 46 chromosomes. This total number of chromosomes is called the chromosome number, when a species has a varying chromosome number, e. g. a diploid and tetraploid form, the chromosome number is called diploid number in the diploid form, and tetraploid number in the tetraploid form. The number of found in a single complete set of chromosomes is called the monoploid number. The haploid number is unique to gametes, and refers to the number of chromosomes found in a gamete. The haploid number for humans is 23, and the monoploid number equals 46 divided by the level of 2. When a human germ cell undergoes meiosis the two sets of 23 chromosomes are split in half to form gametes, after fusion of a male and a female gamete both containing 1 set of 23 chromosomes, the resulting zygote has 46 chromosomes,2 sets of 23 chromosomes. The common potato is an example of an organism, carrying four sets of chromosomes. The potato plant inherits two sets of 12 chromosomes from the parent, and two sets of 12 chromosomes from the ovule parent. The four sets combined provide a complement of 48 chromosomes. The monoploid number equals the number divided by the ploidy level,48 chromosomes in total divided by a ploidy level of 4 equals a monoploid number of 12. Because the chromosome number is reduced only by the specialized process of meiosis. However, in many situations somatic cells double their number by means of endoreduplication as an aspect of cellular differentiation. When a germ cell with an number of chromosomes undergoes meiosis. Triploid organisms for instance are usually sterile, because of this, triploidy is a common way of making seedless fruit such as bananas and watermelons. If the fertilization of human gametes results in 3 sets of chromosomes the condition is called triploid syndrome, the term ploidy is a back-formation from haploidy and diploidy
Diploid
–
Diploid cells have two
homologous copies of each
chromosome.
13.
Glume
–
In botany, a glume is a bract below a spikelet in the inflorescence of grasses or the flowers of sedges. There are two types of bracts in the spikelets of grasses, the lemma and palea. In grasses, two known as glumes form the lowermost organs of a spikelet. Glumes may be similar in form to the lemmas, the bracts at the base of each floret, in sedges, by contrast, a glume is a scale at the base of each flower in a spikelet
Glume
–
Rice chaff
Glume
–
Spikelets of a hulled wheat,
einkorn
14.
Husk
–
Husk in botany is the outer shell or coating of a seed. It often refers to the outer covering of an ear of maize as it grows on the plant. Literally, a husk or hull includes the outer covering of a seed. It can also refer to the exuvia of bugs or small animals left behind after moulting, in cooking, hull can also refer to other waste parts of fruits and vegetables, notably the cap or sepal of a strawberry. The husk of a legume and some similar fruits is called a pod, plantago-seed mucilage is often referred to as husk, or psyllium husk. Crop plants of species have been selected that have hulless seeds, including pumpkins, oats. Husking of corn is the process of removing its outer layers, dehulling is the process of removing the hulls from beans and other seeds. This is sometimes done using a known as a huller. To prepare the seeds to have extracted from them, they are cleaned to remove any foreign objects. Next, the seeds have their hulls, or outer coverings, or husk, there are three different types of dehulling systems that can be used to process soybeans, Hot dehulling, warm dehulling and cold dehulling. Hot dehulling is the system offered in areas where beans are processed directly from the field, warm dehulling is often used by processors who import their soybeans. Cold dehulling is used in plants that have existing drying and conditioning equipment, the different dehulling temperature options are for different types of production, beans and preparation equipment. In third-world countries, husking and dehulling is often done by hand using a large mortar. These are usually made of wood, and operated by one or more people, the husk is biodegradable and may be composted. Chaff Gum Horsebread, a type of bread in which the chaff is not removed Rice hulls Rice pounder Threshing Winnowing
Husk
–
Corn being husked in the yard of a
Dungan farmer in
Kyrgyzstan
Husk
–
Women in
Cape Verde using multiple pestles in a large
mortar
Husk
–
History
Husk
15.
Grain
–
Grains are small, hard, dry seeds, with or without attached hulls or fruit layers, harvested for human or animal consumption. Agronomists also call the plants producing such seeds grain crops, the two main types of commercial grain crops are cereals such as wheat and rye, and legumes such as beans and soybeans. Because of the ubiquity of grain as a source, the term grain is often used to describe something small. After being harvested, dry grains are more durable than other foods, such as starchy fruits. Thus, major global commodity markets exist for canola, maize, rice, soybeans, wheat, in botany, grains and cereals are synonymous with caryopses, the fruits of the grass family. In agronomy and commerce, seeds or fruits from other plant families are called if they resemble caryopses. For example, amaranth is sold as grain amaranth, and amaranth products may be described as whole grains, the pre-Hispanic civilizations of the Andes had grain-based food systems but, at the higher elevations, none of the grains was a cereal. All three grains native to the Andes are broad-leafed plants rather than such as corn, rice. All cereal crops are members of the grass family, cereal grains contain a substantial amount of starch, a carbohydrate that provides dietary energy. As is the case all other whole plant foods, pulses also contain carbohydrate. Vegetable oils provide dietary energy and some essential fatty acids and they are also used as fuel or lubricants. Those who handle grain at grain facilities may encounter numerous occupational hazards, risks include grain entrapment, where workers are submerged in the grain and unable to remove themselves, explosions caused by fine particles of grain dust, and falls
Grain
–
Food grains at a weekly market
Grain
–
Cereal grain seeds from left to right:
wheat,
spelt,
barley,
oat.
Grain
–
Barley
Grain
–
Rye grains.
16.
Pre-Pottery Neolithic B
–
Pre-Pottery Neolithic B is a division of the Neolithic developed by Kathleen Kenyon during her archaeological excavations at Jericho in the West Bank. In addition the flint tool kit of the period is new, one of its major elements is the naviform core. This is the first period in which architectural styles of the southern Levant became primarily rectilinear, earlier typical dwellings were circular, elliptical, pyrotechnology was highly developed in this period. During this period, one of the features of houses is evidenced by a thick layer of white clay plaster floors highly polished. It is believed that the use of plaster for floor. The earliest proto-pottery was White Ware vessels, made from lime and gray ash, built up around baskets before firing, sites from this period found in the Levant utilizing rectangular floor plans and plastered floor techniques were found at Ain Ghazal, Yiftahel, and Abu Hureyra. The period is dated to between ca.10,700 and ca.8,000 BP or 7000 -6000 BCE. Danielle Stordeurs recent work at Tell Aswad, an agricultural village between Mount Hermon and Damascus could not validate Henri de Contensons earlier suggestion of a PPNA Aswadian culture. Instead, they found evidence of a fully established PPNB culture at 8700 BC at Aswad, similar sites to Tell Aswad in the Damascus Basin of the same age were found at Tell Ramad and Tell Ghoraifé. How a PPNB culture could spring up in this location, practicing domesticated farming from 8700 BC has been the subject of speculation. Like the earlier PPNA people, the PPNB culture developed from the Earlier Natufian but shows evidence of a northerly origin, work at the site of Ain Ghazal in Jordan has indicated a later Pre-Pottery Neolithic C period which existed between 8,200 and 7,900 BP. Cultures practicing this lifestyle spread down the Red Sea shoreline and moved east from Syria into southern Iraq
Pre-Pottery Neolithic B
–
Geography
Pre-Pottery Neolithic B
–
Pre-Pottery Neolithic B
17.
Turkey
–
Turkey, officially the Republic of Turkey, is a transcontinental country in Eurasia, mainly in Anatolia in Western Asia, with a smaller portion on the Balkan peninsula in Southeast Europe. Turkey is a democratic, secular, unitary, parliamentary republic with a cultural heritage. The country is encircled by seas on three sides, the Aegean Sea is to the west, the Black Sea to the north, and the Mediterranean Sea to the south. The Bosphorus, the Sea of Marmara, and the Dardanelles, Ankara is the capital while Istanbul is the countrys largest city and main cultural and commercial centre. Approximately 70-80% of the countrys citizens identify themselves as ethnic Turks, other ethnic groups include legally recognised and unrecognised minorities. Kurds are the largest ethnic minority group, making up approximately 20% of the population, the area of Turkey has been inhabited since the Paleolithic by various ancient Anatolian civilisations, as well as Assyrians, Greeks, Thracians, Phrygians, Urartians and Armenians. After Alexander the Greats conquest, the area was Hellenized, a process continued under the Roman Empire. The Seljuk Sultanate of Rûm ruled Anatolia until the Mongol invasion in 1243, the empire reached the peak of its power in the 16th century, especially during the reign of Suleiman the Magnificent. During the war, the Ottoman government committed genocides against its Armenian, Assyrian, following the war, the conglomeration of territories and peoples that formerly comprised the Ottoman Empire was partitioned into several new states. Turkey is a member of the UN, an early member of NATO. Turkeys growing economy and diplomatic initiatives have led to its recognition as a regional power while her location has given it geopolitical, the name of Turkey is based on the ethnonym Türk. The first recorded use of the term Türk or Türük as an autonym is contained in the Old Turkic inscriptions of the Göktürks of Central Asia, the English name Turkey first appeared in the late 14th century and is derived from Medieval Latin Turchia. Similarly, the medieval Khazar Empire, a Turkic state on the shores of the Black. The medieval Arabs referred to the Mamluk Sultanate as al-Dawla al-Turkiyya, the Ottoman Empire was sometimes referred to as Turkey or the Turkish Empire among its European contemporaries. The Anatolian peninsula, comprising most of modern Turkey, is one of the oldest permanently settled regions in the world, various ancient Anatolian populations have lived in Anatolia, from at least the Neolithic period until the Hellenistic period. Many of these peoples spoke the Anatolian languages, a branch of the larger Indo-European language family, in fact, given the antiquity of the Indo-European Hittite and Luwian languages, some scholars have proposed Anatolia as the hypothetical centre from which the Indo-European languages radiated. The European part of Turkey, called Eastern Thrace, has also been inhabited since at least forty years ago. It is the largest and best-preserved Neolithic site found to date, the settlement of Troy started in the Neolithic Age and continued into the Iron Age
Turkey
–
Some
henges at
Göbekli Tepe were erected as far back as 12,000 BC, predating those of
Stonehenge,
England by almost ten millennia.
Turkey
–
Flag
Turkey
–
The Lion Gate in
Hattusa, capital of the
Hittite Empire. The city's history dates back to the 6th millennium BC.
Turkey
–
The
Library of Celsus in
Ephesus was built by the
Romans in 135 AD. The
Temple of Artemis in Ephesus, built by king
Croesus of
Lydia in the 6th century BC, was one of the
Seven Wonders of the Ancient World.
18.
Fertile Crescent
–
The Fertile Crescent is a crescent-shaped region containing the comparatively moist and fertile land of otherwise arid and semi-arid Western Asia, the Nile Valley and Nile Delta. Having originated in the study of ancient history, the concept developed and today retains meanings in international geopolitics. In current usage, all definitions of the Fertile Crescent include Mesopotamia, the land in and around the Tigris and Euphrates rivers, and the Levant, the eastern coast of the Mediterranean Sea. The region saw the development of some of the earliest human civilizations, technological advances made in the region include the development of writing, glass, the wheel, agriculture, and the use of irrigation. It is a region consisting chiefly of mountains in the north, the end of the western wing is Palestine, Assyria makes up a large part of the center, while the end of the eastern wing is Babylonia. This great semicircle, for lack of a name, may be called the Fertile Crescent and this desert-bay is a limestone plateau of some height—too high indeed to be watered by the Tigris and Euphrates, which have cut cañons obliquely across it. Hence we are obliged to coin a term and call it the Fertile Crescent, water sources include the Jordan River. The inner boundary is delimited by the dry climate of the Syrian Desert to the south, around the outer boundary are the Anatolian highlands to the north and the Sahara Desert to the west. As crucial as rivers and marshlands were to the rise of civilization in the Fertile Crescent, the area is important as the bridge between Africa and Eurasia. Coupled with the Saharan pump theory, this Middle Eastern land-bridge is of importance to the modern distribution of Old World flora and fauna. The Fertile Crescent had many diverse climates, and major climatic changes encouraged the evolution of many r type annual plants, the regions dramatic variety of elevation gave rise to many species of edible plants for early experiments in cultivation. The Fertile Crescent has a record of past human activity. The western zone around the Jordan and upper Euphrates rivers gave rise to the first known Neolithic farming settlements and this region, alongside Mesopotamia, also saw the emergence of early complex societies during the succeeding Bronze Age. There is also evidence from the region for writing and the formation of hierarchical statelevel societies. This has earned the region the nickname The cradle of civilization and it is in this region where the first libraries appeared, some 5,000 years ago. The oldest known library was found in northern Syria, in the ruins of Ebla— a major commercial center that was destroyed around 1650 BCE, both the Tigris and Euphrates start in the Taurus Mountains of what is today Turkey. Farmers in southern Mesopotamia had to protect their fields from flooding each year, to protect against flooding, they made levees. Since the Bronze Age, the regions natural fertility has been extended by irrigation works
Fertile Crescent
–
The Fertile Crescent at maximum defined extent, with the names of ancient civilizations found there.
19.
Anatolia
–
Anatolia, in geography known as Asia Minor, Asian Turkey, Anatolian peninsula, or Anatolian plateau, is the westernmost protrusion of Asia, which makes up the majority of modern-day Turkey. The region is bounded by the Black Sea to the north, the Mediterranean Sea to the south, the Sea of Marmara forms a connection between the Black and Aegean Seas through the Bosphorus and Dardanelles straits and separates Anatolia from Thrace on the European mainland. Traditionally, Anatolia is considered to extend in the east to a line between the Gulf of Alexandretta and the Black Sea to the Armenian Highlands, thus, traditionally Anatolia is the territory that comprises approximately the western two-thirds of the Asian part of Turkey. The Turkification of Anatolia began under the Seljuk Empire in the late 11th century, however, various non-Turkic languages continue to be spoken by minorities in Anatolia today, including Kurdish, Assyrian, Armenian, Arabic, Laz, Georgian, and Greek. Traditionally, Anatolia is considered to extend in the east to a line running from the Gulf of Alexandretta to the Black Sea. This traditional geographical definition is used, for example, in the latest edition of Merriam-Websters Geographical Dictionary, under this definition, Anatolia is bounded to the east by the Armenian Highlands, and the Euphrates before that river bends to the southeast to enter Mesopotamia. To the southeast, it is bounded by the ranges that separate it from the Orontes valley in Syria, the first name the Greeks used for the Anatolian peninsula was Ἀσία, presumably after the name of the Assuwa league in western Anatolia. As the name of Asia came to be extended to areas east of the Mediterranean. The name Anatolia derives from the Greek ἀνατολή meaning “the East” or more literally “sunrise”, the precise reference of this term has varied over time, perhaps originally referring to the Aeolian, Ionian and Dorian colonies on the west coast of Asia Minor. In the Byzantine Empire, the Anatolic Theme was a theme covering the western, the modern Turkish form of Anatolia is Anadolu, which again derives from the Greek name Aνατολή. The Russian male name Anatoly and the French Anatole share the same linguistic origin, in English the name of Turkey for ancient Anatolia first appeared c. It is derived from the Medieval Latin Turchia, which was used by the Europeans to define the Seljuk controlled parts of Anatolia after the Battle of Manzikert. Human habitation in Anatolia dates back to the Paleolithic, neolithic Anatolia has been proposed as the homeland of the Indo-European language family, although linguists tend to favour a later origin in the steppes north of the Black Sea. However, it is clear that the Anatolian languages, the oldest branch of Indo-European, have spoken in Anatolia since at least the 19th century BC. The earliest historical records of Anatolia stem from the southeast of the region and are from the Mesopotamian-based Akkadian Empire during the reign of Sargon of Akkad in the 24th century BC, scholars generally believe the earliest indigenous populations of Anatolia were the Hattians and Hurrians. The region was famous for exporting raw materials, and areas of Hattian-, one of the numerous cuneiform records dated circa 20th century BC, found in Anatolia at the Assyrian colony of Kanesh, uses an advanced system of trading computations and credit lines. They were speakers of an Indo-European language, the Hittite language, originating from Nesa, they conquered Hattusa in the 18th century BC, imposing themselves over Hattian- and Hurrian-speaking populations. According to the most widely accepted Kurgan theory on the Proto-Indo-European homeland, however, the Hittites adopted the cuneiform script, invented in Mesopotamia
Anatolia
–
The traditional definition of Anatolia within modern Turkey
Anatolia
–
The location of
Turkey (within the rectangle) in reference to the
European continent. Anatolia roughly corresponds to the
Asian part of Turkey, except the eastern parts historically known as the
Armenian Highlands
Anatolia
–
1907 map of Asia Minor, showing the local ancient kingdoms
Anatolia
–
Mural of
aurochs, a
deer, and humans in
Çatalhöyük, which is the largest and best-preserved Neolithic site found to date. It was registered as a
UNESCO World Heritage Site in 2012.
20.
Balkans
–
The Balkan Peninsula, or the Balkans, is a peninsula and a cultural area in Eastern and Southeastern Europe with various and disputed borders. The region takes its name from the Balkan Mountains that stretch from the Serbia-Bulgaria border to the Black Sea, the highest point of the Balkans is Mount Musala 2,925 metres in the Rila mountain range. In Turkish, Balkan means a chain of wooded mountains, the name is still preserved in Central Asia with the Balkan Daglary and the Balkan Province of Turkmenistan. A less popular hypothesis regarding its etymology is that it derived from the Persian Balā-Khāna, from Antiquity through the Middle Ages, the Balkan Mountains had been called by the local Thracian name Haemus. According to Greek mythology, the Thracian king Haemus was turned into a mountain by Zeus as a punishment, a reverse name scheme has also been suggested. D. Dechev considers that Haemus is derived from a Thracian word *saimon, a third possibility is that Haemus derives from the Greek word haema meaning blood. The myth relates to a fight between Zeus and the monster/titan Typhon, Zeus injured Typhon with a thunder bolt and Typhons blood fell on the mountains, from which they got their name. The earliest mention of the name appears in an early 14th-century Arab map, the Ottomans first mention it in a document dated from 1565. There has been no other documented usage of the word to refer to the region before that, there is also a claim about an earlier Bulgar Turkic origin of the word popular in Bulgaria, however it is only an unscholarly assertion. The word was used by the Ottomans in Rumelia in its meaning of mountain, as in Kod̲j̲a-Balkan, Čatal-Balkan, and Ungurus-Balkani̊. The concept of the Balkans was created by the German geographer August Zeune in 1808, during the 1820s, Balkan became the preferred although not yet exclusive term alongside Haemus among British travelers. Among Russian travelers not so burdened by classical toponymy, Balkan was the preferred term, zeunes goal was to have a geographical parallel term to the Italic and Iberian Peninsula, and seemingly nothing more. The gradually acquired political connotations are newer and, to a large extent, after the dissolution of Yugoslavia beginning in June 1991, the term Balkans again received a negative meaning, especially in Croatia and Slovenia, even in casual usage. A European Union initiative of 1999 is called the Stability Pact for South Eastern Europe, and its northern boundary is often given as the Danube, Sava and Kupa Rivers. The Balkan Peninsula has an area of about 470,000 km2. It is more or less identical to the known as Southeastern Europe. As of 1920 until World War II, Italy included Istria, the current territory of Italy includes only the small area around Trieste inside the Balkan Peninsula. However, the regions of Trieste and Istria are not usually considered part of the Balkans by Italian geographers, the Western Balkans is a neologism coined to describe the countries of ex-Yugoslavia and Albania
Balkans
–
The Balkan Peninsula, as defined by the
Danube -
Sava -
Kupa line
Balkans
–
The Peninsula's most extensive definition, bordered by water on three sides and connected with a line on the fourth
Balkans
–
Panorama of
Stara Planina. Its highest peak is
Botev at a height of 2,376 m.
Balkans
–
View toward
Rila, the highest mountain in the Balkans which reaches 2925 m
21.
Dead Sea
–
The Dead Sea, is a salt lake bordered by Jordan to the east and Israel and Palestine to the west. Its surface and shores are 430.5 metres below sea level, the Dead Sea is 304 m deep, the deepest hypersaline lake in the world. With 34. 2% salinity, it is 9.6 times as salty as the ocean and this salinity makes for a harsh environment in which plants and animals cannot flourish, hence its name. The Dead Sea is 50 kilometres long and 15 kilometres wide at its widest point and it lies in the Jordan Rift Valley and its main tributary is the Jordan River. The Dead Sea has attracted visitors from around the Mediterranean basin for thousands of years and it was one of the worlds first health resorts, and it has been the supplier of a wide variety of products, from asphalt for Egyptian mummification to potash for fertilizers. People also use the salt and the minerals from the Dead Sea to create cosmetics, the Dead Sea water has a density of 1.24 kg/litre, which makes swimming similar to floating. The Dead Sea is receding at an alarming rate, multiple canals and pipelines were proposed to reduce its recession, which had begun causing many problems. The Red Sea–Dead Sea Water Conveyance project, carried out by Jordan, will provide water to neighboring countries, the first phase of the project is scheduled to begin in 2018 and be completed in 2021. In Hebrew, the Dead Sea is Yām ha-Melaḥ, meaning sea of salt, the Bible uses this term alongside two others, the Sea of the Arabah, and the Eastern Sea. The designation Dead Sea never appears in the Bible, in prose sometimes the term Yām ha-Māvet is used, due to the scarcity of aquatic life there. In Arabic the Dead Sea is called al-Bahr al-Mayyit, or less commonly baḥrᵘ lūṭᵃ, another historic name in Arabic was the Sea of Zoʼar, after a nearby town in biblical times. The Greeks called it Lake Asphaltites, the Dead Sea is an endorheic lake located in the Jordan Rift Valley, a geographic feature formed by the Dead Sea Transform. This left lateral-moving transform fault lies along the plate boundary between the African Plate and the Arabian Plate. It runs between the East Anatolian Fault zone in Turkey and the end of the Red Sea Rift offshore of the southern tip of Sinai. It is here that the Upper Jordan River/Sea of Galilee/Lower Jordan River water system comes to an end. The Jordan River is the major water source flowing into the Dead Sea, although there are small perennial springs under and around the Dead Sea, forming pools. The Mujib River, biblical Arnon, is one the larger sources of the Dead Sea other than the Jordan. The Wadi Mujib valley,420 m below the sea level in the southern of Jordan valley, is a biosphere reserve, other more substantial sources are Wadi Darajeh /Nahal Dragot, and Nahal Arugot
Dead Sea
–
A view of the sea from the shore
Dead Sea
–
Satellite photograph showing the location of the Dead Sea
Dead Sea
–
Pebbles cemented with
halite on the western shore of the Dead Sea near Ein Gedi.
Dead Sea
–
Halite deposits (and teepee structure) along the western Dead Sea coast.
22.
Mutation
–
In biology, a mutation is the permanent alteration of the nucleotide sequence of the genome of an organism, virus, or extrachromosomal DNA or other genetic elements. Mutations may also result from insertion or deletion of segments of DNA due to mobile genetic elements, mutations may or may not produce discernible changes in the observable characteristics of an organism. Mutations play a part in normal and abnormal biological processes including, evolution, cancer, and the development of the immune system. The genomes of RNA viruses are based on RNA rather than DNA, the RNA viral genome can be double stranded or single stranded. In some of these viruses replication occurs quickly and there are no mechanisms to check the genome for accuracy and this error-prone process often results in mutations. Mutation can result in different types of change in sequences. Mutations in genes can either have no effect, alter the product of a gene, mutations can also occur in nongenic regions. Mutations can involve the duplication of large sections of DNA, usually through genetic recombination and these duplications are a major source of raw material for evolving new genes, with tens to hundreds of genes duplicated in animal genomes every million years. Most genes belong to larger families of shared ancestry, known as homology. Here, protein domains act as modules, each with a particular and independent function, that can be mixed together to produce genes encoding new proteins with novel properties. For example, the eye uses four genes to make structures that sense light. Another advantage of duplicating a gene is that this increases engineering redundancy, other types of mutation occasionally create new genes from previously noncoding DNA. Changes in chromosome number may involve even larger mutations, where segments of the DNA within chromosomes break and then rearrange. For example, in the Homininae, two fused to produce human chromosome 2, this fusion did not occur in the lineage of the other apes. Sequences of DNA that can move about the genome, such as transposons, make up a fraction of the genetic material of plants and animals. For example, more than a million copies of the Alu sequence are present in the genome. Another effect of these mobile DNA sequences is that when they move within a genome, they can mutate or delete existing genes, nonlethal mutations accumulate within the gene pool and increase the amount of genetic variation. The abundance of some changes within the gene pool can be reduced by natural selection, while other more favorable mutations may accumulate
Mutation
–
A mutation has caused this garden
moss rose to produce flowers of different colors. This is a
somatic mutation that may also be passed on in the
germline.
Mutation
–
Diagrammatic representation of the
divergence of modern
taxonomic groups from their
common ancestor
23.
Geographic coordinate system
–
A geographic coordinate system is a coordinate system used in geography that enables every location on Earth to be specified by a set of numbers, letters or symbols. The coordinates are chosen such that one of the numbers represents a vertical position. A common choice of coordinates is latitude, longitude and elevation, to specify a location on a two-dimensional map requires a map projection. The invention of a coordinate system is generally credited to Eratosthenes of Cyrene. Ptolemy credited him with the adoption of longitude and latitude. Ptolemys 2nd-century Geography used the prime meridian but measured latitude from the equator instead. Mathematical cartography resumed in Europe following Maximus Planudes recovery of Ptolemys text a little before 1300, in 1884, the United States hosted the International Meridian Conference, attended by representatives from twenty-five nations. Twenty-two of them agreed to adopt the longitude of the Royal Observatory in Greenwich, the Dominican Republic voted against the motion, while France and Brazil abstained. France adopted Greenwich Mean Time in place of local determinations by the Paris Observatory in 1911, the latitude of a point on Earths surface is the angle between the equatorial plane and the straight line that passes through that point and through the center of the Earth. Lines joining points of the same latitude trace circles on the surface of Earth called parallels, as they are parallel to the equator, the north pole is 90° N, the south pole is 90° S. The 0° parallel of latitude is designated the equator, the plane of all geographic coordinate systems. The equator divides the globe into Northern and Southern Hemispheres, the longitude of a point on Earths surface is the angle east or west of a reference meridian to another meridian that passes through that point. All meridians are halves of great ellipses, which converge at the north and south poles, the prime meridian determines the proper Eastern and Western Hemispheres, although maps often divide these hemispheres further west in order to keep the Old World on a single side. The antipodal meridian of Greenwich is both 180°W and 180°E, the combination of these two components specifies the position of any location on the surface of Earth, without consideration of altitude or depth. The grid formed by lines of latitude and longitude is known as a graticule, the origin/zero point of this system is located in the Gulf of Guinea about 625 km south of Tema, Ghana. To completely specify a location of a feature on, in, or above Earth. Earth is not a sphere, but a shape approximating a biaxial ellipsoid. It is nearly spherical, but has an equatorial bulge making the radius at the equator about 0. 3% larger than the radius measured through the poles, the shorter axis approximately coincides with the axis of rotation
Geographic coordinate system
–
Longitude lines are perpendicular and latitude lines are parallel to the equator.
24.
Emmer
–
Emmer wheat, also known as farro especially in Italy, or hulled wheat, is a type of awned wheat. The domesticated species are Triticum turgidum subsp, dicoccum and Triticum turgidum conv. durum. The wild species is called Triticum turgidum subsp, along with einkorn wheat, emmer was one of the first crops domesticated in the Near East. It was widely cultivated in the ancient world, but is now a relict crop in regions of Europe. Strong similarities in morphology and genetics show that wild emmer is the wild ancestor, because wild and domesticated emmer are interfertile with other tetraploid wheats, some taxonomists consider all tetraploid wheats to belong to one species, T. turgidum. Under this scheme, the two forms are recognized at subspecies level, thus T. turgidum subsp, either naming system is equally valid, the latter lays more emphasis on genetic similarities. For a wider discussion, see Wheat#Genetics & Breeding and Wheat taxonomy Wild emmer grows wild in the Near East, like einkorn and spelt wheats, emmer is a hulled wheat. In other words, it has strong glumes that enclose the grains, on threshing, a hulled wheat spike breaks up into spikelets. These require milling or pounding to release the grains from the glumes, Wild emmer wheat spikelets effectively self-cultivate by propelling themselves mechanically into soils with their awns. During a period of increased humidity during the night, the awns of the spikelet become erect and draw together, and in the process push the grain into the soil. During the daytime, the humidity drops and the awns slacken back again, however, fine hairs on the awns act as hooks in the soil. During the course of alternating stages of daytime and nighttime humidity, the awns pumping movements, Wild emmer is native to the Fertile Crescent of the Middle East, growing in the grass and woodland of hill country from modern-day Israel to Iran. The origin of wild emmer has been suggested, without universal agreement among scholars, in 1906, Aaron Aaronsohns discovery of wild emmer wheat growing in Rosh Pinna created a stir in the botanical world. Emmer wheat has been found in excavations and ancient tombs. Emmer was collected from the wild and eaten by hunter gatherers for thousands of years before its domestication. Grains of wild emmer discovered at Ohalo II had a radiocarbon dating of 17,000 BC and at the Pre Pottery Neolithic A site of Netiv Hagdud are 10, the location of the earliest site of emmer domestication is still unclear and under debate. Emmer is found in a number of Neolithic sites scattered around the fertile crescent. From its earliest days of cultivation, emmer was a more prominent crop than its contemporaries and competitors, einkorn wheat
Emmer
–
Emmer wheat
Emmer
–
History
25.
Epi-Paleolithic
–
Epipaleolithic is a term used for the final Upper Palaeolithic industries occurring at the end of the final glaciation which appear to merge technologically into the Mesolithic. The period is dated from c. 20,000 BP to c. 10,500 BP. The term is used as a synonym of Mesolithic. Inversely, those that are in transitional ways towards artificial production of food are inscribed in the Mesolithic, epipalaeolithic hunter-gatherers, generally nomadic, made relatively advanced tools from small flint or obsidian blades, known as microliths, that were hafted in wooden implements. The Epipaleolithic is best understood when discussing the southern Levant, as the period is well documented due to preservation at the site. Of these animals, it is likely only the equids were migrational
Epi-Paleolithic
–
Azilian points,
microliths from epipaleolithic northern Spain and southern France.
26.
Pre-Pottery Neolithic A
–
Pre-Pottery Neolithic A denotes the first stage in early Levantine and Anatolian Neolithic culture, dating c. 11,500 – c. 10,000 BP. Archaeological remains are located in the Levantine and Upper Mesopotamian region of the Fertile Crescent, the Pre-Pottery Neolithic A and the following Pre-Pottery Neolithic B were originally defined by Kathleen Kenyon in the type site of Jericho. During this time, pottery was not yet in use, PPNA succeeds the Natufian culture of the Epipaleolithic. PPNA archaeological sites are larger than those of the preceding Natufian hunter-gatherer culture. PPNA settlements are characterized by round, semi-subterranean houses with stone foundations, the upper walls were constructed of unbaked clay mudbricks with plano-convex cross-sections. The hearths were small, and covered with cobbles, heated rocks were used in cooking, which led to an accumulation of fire-cracked rock in the buildings, and almost every settlement contained storage bins made of either stones or mud-brick. One of the most notable PPNA settlements is Jericho, thought to be the worlds first town, the PPNA town contained a population of up to 2, 000–3,000 people, and was protected by a massive stone wall and tower. There is much debate over the function of the wall, for there is no evidence of any serious warfare at this time, one possibility is the wall was built to protect the salt resources of Jericho. PPNA cultures are unique for their practices, and Kenyon characterized them as living with their dead. Kenyon found no fewer than 279 burials, below floors, under household foundations, in the PPNB period, skulls were often dug up and reburied, or mottled with clay and displayed. The lithic industry is based on blades struck from regular cores, sickle-blades and arrowheads continue traditions from the late Natufian culture, transverse-blow axes and polished adzes appear for the first time. Sedentism of this allowed for the cultivation of local grains, such as barley and wild oats. Sites such as Dhra′ and Jericho retained a hunting lifestyle until the PPNB period and this period of cultivation is considered pre-domestication, but may have begun to develop plant species into the domesticated forms they are today. Deliberate, extended-period storage was possible by the use of suspended floors for air circulation and protection from rodents. This practice precedes the emergence of domestication and large-scale sedentary communities by at least 1,000 years, granaries are positioned in places between other buildings early on c. 11,500 BP. However, beginning around 10,500 BP, they were moved inside houses and this change might reflect changing systems of ownership and property as granaries shifted from a communal use and ownership to become under the control of households or individuals. Moreover, uilding granaries may have been the most important feature in increasing sedentism that required active community participation in new life-ways. With more sites becoming known, archaeologists have defined a number of variants, Sultanian in the Jordan River valley
Pre-Pottery Neolithic A
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El Khiam point, schematic drawing
Pre-Pottery Neolithic A
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Pre-Pottery Neolithic A
Pre-Pottery Neolithic A
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Geography
27.
Caucasus
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The Caucasus /ˈkɔːkəsəs/ or Caucasia /kɔːˈkeɪʒə/ is a region at the border of Europe and Asia, situated between the Black and the Caspian seas. It is home to the Caucasus Mountains, which contain Europes highest mountain, politically, the Caucasus region is separated between northern and southern parts. The southern parts consist of independent sovereign states, and the parts are under the jurisdiction of the Russian Federation. The region is known for its diversity, aside from Indo-European and Turkic languages, the Kartvelian, Northwest Caucasian. Pliny the Elders Natural History derives the name of the Caucasus from Scythian kroy-khasis, German linguist Paul Kretschmer notes that the Latvian word Kruvesis also means ice. According to German philologists Otto Schrader and Alfons A. Nehring, the South Caucasus region and southern Dagestan were the furthest points of Persian expansions, with areas to the north of Caucasus Mountains practically impregnable. The mythological mountain of Qaf, the worlds highest mountain that ancient lore shrouded in mystery, was said to be situated in this region, therefore, the Caucasus might be associated with the legendary mountain. The Ciscaucasus contains the majority of the Greater Caucasus Mountain range. It includes Southwestern Russia and northern parts of Georgia and Azerbaijan, the Transcaucasus is bordered on the north by Russia, on the west by the Black Sea and Turkey, on the east by the Caspian Sea, and on the south by Iran. It includes the Caucasus Mountains and surrounding lowlands, all of Armenia, Azerbaijan and Georgia are in South Caucasus. The main Greater Caucasus range is generally perceived to be the line between Asia and Europe. The highest peak in the Caucasus is Mount Elbrus in the western Ciscaucasus in Russia, the Caucasus is one of the most linguistically and culturally diverse regions on Earth. The nation states that comprise the Caucasus today are the post-Soviet states Georgia, Armenia, three territories in the region claim independence but are recognized as such by only a handful or by no independent states, Nagorno-Karabakh, Abkhazia and South Ossetia. Abkhazia and South Ossetia are recognised by the majority of independent states as part of Georgia, the Russian divisions include Krasnodar Krai, Stavropol Krai, and the autonomous republics of Adygea, Karachay–Cherkessia, Kabardino-Balkaria, North Ossetia, Ingushetia, Chechnya, and Dagestan. The region has many different languages and language families, there are more than 50 ethnic groups living in the region. Russian is used as a common language, today the peoples of the Northern and Southern Caucasus tend to be either Eastern Orthodox Christians, Oriental Orthodox Christians, or Sunni Muslims. Shia Islam has had many adherents historically in Azerbaijan, located in the part of the region. Located on the peripheries of Turkey, Iran, and Russia, the region has been an arena for political, military, religious, throughout its history, the Caucasus was usually incorporated into the Iranian world
Caucasus
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Caucasus from the sea by
Ivan Aivazovsky, 1899
Caucasus
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Caucasus mountain range
Caucasus
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View of the Caucasus Mountains in
Dagestan, Russia
Caucasus
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Petroglyphs in
Gobustan,
Azerbaijan, dating back to 10,000 BC indicating a thriving culture. It is a
UNESCO World Heritage Site.
28.
Emmer wheat
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Emmer wheat, also known as farro especially in Italy, or hulled wheat, is a type of awned wheat. The domesticated species are Triticum turgidum subsp, dicoccum and Triticum turgidum conv. durum. The wild species is called Triticum turgidum subsp, along with einkorn wheat, emmer was one of the first crops domesticated in the Near East. It was widely cultivated in the ancient world, but is now a relict crop in regions of Europe. Strong similarities in morphology and genetics show that wild emmer is the wild ancestor, because wild and domesticated emmer are interfertile with other tetraploid wheats, some taxonomists consider all tetraploid wheats to belong to one species, T. turgidum. Under this scheme, the two forms are recognized at subspecies level, thus T. turgidum subsp, either naming system is equally valid, the latter lays more emphasis on genetic similarities. For a wider discussion, see Wheat#Genetics & Breeding and Wheat taxonomy Wild emmer grows wild in the Near East, like einkorn and spelt wheats, emmer is a hulled wheat. In other words, it has strong glumes that enclose the grains, on threshing, a hulled wheat spike breaks up into spikelets. These require milling or pounding to release the grains from the glumes, Wild emmer wheat spikelets effectively self-cultivate by propelling themselves mechanically into soils with their awns. During a period of increased humidity during the night, the awns of the spikelet become erect and draw together, and in the process push the grain into the soil. During the daytime, the humidity drops and the awns slacken back again, however, fine hairs on the awns act as hooks in the soil. During the course of alternating stages of daytime and nighttime humidity, the awns pumping movements, Wild emmer is native to the Fertile Crescent of the Middle East, growing in the grass and woodland of hill country from modern-day Israel to Iran. The origin of wild emmer has been suggested, without universal agreement among scholars, in 1906, Aaron Aaronsohns discovery of wild emmer wheat growing in Rosh Pinna created a stir in the botanical world. Emmer wheat has been found in excavations and ancient tombs. Emmer was collected from the wild and eaten by hunter gatherers for thousands of years before its domestication. Grains of wild emmer discovered at Ohalo II had a radiocarbon dating of 17,000 BC and at the Pre Pottery Neolithic A site of Netiv Hagdud are 10, the location of the earliest site of emmer domestication is still unclear and under debate. Emmer is found in a number of Neolithic sites scattered around the fertile crescent. From its earliest days of cultivation, emmer was a more prominent crop than its contemporaries and competitors, einkorn wheat
Emmer wheat
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Emmer wheat
Emmer wheat
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History
29.
Italy
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Italy, officially the Italian Republic, is a unitary parliamentary republic in Europe. Located in the heart of the Mediterranean Sea, Italy shares open land borders with France, Switzerland, Austria, Slovenia, San Marino, Italy covers an area of 301,338 km2 and has a largely temperate seasonal and Mediterranean climate. Due to its shape, it is referred to in Italy as lo Stivale. With 61 million inhabitants, it is the fourth most populous EU member state, the Italic tribe known as the Latins formed the Roman Kingdom, which eventually became a republic that conquered and assimilated other nearby civilisations. The legacy of the Roman Empire is widespread and can be observed in the distribution of civilian law, republican governments, Christianity. The Renaissance began in Italy and spread to the rest of Europe, bringing a renewed interest in humanism, science, exploration, Italian culture flourished at this time, producing famous scholars, artists and polymaths such as Leonardo da Vinci, Galileo, Michelangelo and Machiavelli. The weakened sovereigns soon fell victim to conquest by European powers such as France, Spain and Austria. Despite being one of the victors in World War I, Italy entered a period of economic crisis and social turmoil. The subsequent participation in World War II on the Axis side ended in defeat, economic destruction. Today, Italy has the third largest economy in the Eurozone and it has a very high level of human development and is ranked sixth in the world for life expectancy. The country plays a prominent role in regional and global economic, military, cultural and diplomatic affairs, as a reflection of its cultural wealth, Italy is home to 51 World Heritage Sites, the most in the world, and is the fifth most visited country. The assumptions on the etymology of the name Italia are very numerous, according to one of the more common explanations, the term Italia, from Latin, Italia, was borrowed through Greek from the Oscan Víteliú, meaning land of young cattle. The bull was a symbol of the southern Italic tribes and was often depicted goring the Roman wolf as a defiant symbol of free Italy during the Social War. Greek historian Dionysius of Halicarnassus states this account together with the legend that Italy was named after Italus, mentioned also by Aristotle and Thucydides. The name Italia originally applied only to a part of what is now Southern Italy – according to Antiochus of Syracuse, but by his time Oenotria and Italy had become synonymous, and the name also applied to most of Lucania as well. The Greeks gradually came to apply the name Italia to a larger region, excavations throughout Italy revealed a Neanderthal presence dating back to the Palaeolithic period, some 200,000 years ago, modern Humans arrived about 40,000 years ago. Other ancient Italian peoples of undetermined language families but of possible origins include the Rhaetian people and Cammuni. Also the Phoenicians established colonies on the coasts of Sardinia and Sicily, the Roman legacy has deeply influenced the Western civilisation, shaping most of the modern world
Italy
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The
Colosseum in Rome, built c. 70 – 80 AD, is considered one of the greatest works of
architecture and
engineering of ancient history.
Italy
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Flag
Italy
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The
Iron Crown of Lombardy, for centuries symbol of the
Kings of Italy.
Italy
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Castel del Monte, built by German Emperor
Frederick II,
UNESCO World Heritage site
. PMID 24253052. doi:10.3390/nu5114553.
