1.
Archaeopteryx
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Archaeopteryx, sometimes referred to by its German name Urvogel, is a genus of bird-like dinosaurs that is transitional between non-avian feathered dinosaurs and modern birds. The name derives from the ancient Greek ἀρχαῖος meaning ancient, and πτέρυξ, between the late nineteenth century and the early twenty-first century, Archaeopteryx had been generally accepted by palaeontologists and popular reference books as the oldest known bird. Older potential avialans have since been identified, including Anchiornis, Xiaotingia, similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, the largest species of Archaeopteryx could grow to about 0.5 m in length. Despite their small size, broad wings, and inferred ability to fly or glide and these features make Archaeopteryx a clear candidate for a transitional fossil between non-avian dinosaurs and birds. Thus, Archaeopteryx plays an important role, not only in the study of the origin of birds and it was named from a single feather in 1861. That same year, the first complete specimen of Archaeopteryx was announced, over the years, ten more fossils of Archaeopteryx have surfaced. Despite variation among these fossils, most experts regard all the remains that have been discovered as belonging to a single species, most of these eleven fossils include impressions of feathers. Because these feathers are of a form, these fossils are evidence that the evolution of feathers began before the Late Jurassic. The type specimen of Archaeopteryx was discovered just two years after Charles Darwin published On the Origin of Species, Archaeopteryx was roughly the size of a raven, with broad wings that were rounded at the ends and a long tail compared to its body length. It could reach up to 500 millimetres in length, with an estimated mass of 0.8 to 1 kilogram. Archaeopteryx feathers, although less documented than its other features, were similar in structure to modern-day bird feathers. Despite the presence of numerous features, Archaeopteryx had many non-avian theropod dinosaur characteristics. Unlike modern birds, Archaeopteryx had small teeth, as well as a bony tail. Because it displays features common to both birds and non-avian dinosaurs, Archaeopteryx has often considered a link between them. For instance, it has an ascending process of the ankle bone, interdental plates, an obturator process of the ischium. In particular, Ostrom found that Archaeopteryx was remarkably similar to the theropod family Dromaeosauridae, specimens of Archaeopteryx were most notable for their well-developed flight feathers. They were markedly asymmetrical and showed the structure of feathers in modern birds. The tail feathers were asymmetrical, again in line with the situation in modern birds
2.
Palaeontological Museum, Munich
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The Paläontologische Museum München is a German national natural history museum situated in Munich, Bavaria. It is associated with the Ludwig-Maximilians-Universität and it has a large collection of fossils of animals and plants such as Mesozoic reptiles, early elephants and saber-toothed cats. One of its highlights is the specimen of the early bird Archaeopteryx discovered in 1991, the museum is also interesting because of the architecture of its building, the former urban college of arts and crafts. List of museums in Germany List of natural history museums Media related to Paläontologisches Museum München at Wikimedia Commons Homepage Munich museums
3.
Jurassic
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The Jurassic is a geologic period and system that spans 56.3 million years from the end of the Triassic Period 201.3 million years ago to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles, the start of the period is marked by the major Triassic–Jurassic extinction event. The Jurassic is named after the Jura Mountains within the European Alps, by the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses, Laurasia to the north and Gondwana to the south. This created more coastlines and shifted the continental climate from dry to humid, on land, the fauna transitioned from the Triassic fauna, dominated by both dinosauromorph and crocodylomorph archosaurs, to one dominated by dinosaurs alone. The first birds also appeared during the Jurassic, having evolved from a branch of theropod dinosaurs, other major events include the appearance of the earliest lizards, and the evolution of therian mammals, including primitive placentals. Crocodilians made the transition from a terrestrial to a mode of life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, the chronostratigraphic term Jurassic is directly linked to the Jura Mountains. The name Jura is derived from the Celtic root jor, which was Latinised into juria, the Jurassic period is divided into the Early Jurassic, Middle, and Late Jurassic epochs. The Jurassic System, in stratigraphy, is divided into the Lower Jurassic, Middle, the separation of the term Jurassic into three sections goes back to Leopold von Buch. The Jurassic North Atlantic Ocean was relatively narrow, while the South Atlantic did not open until the following Cretaceous period, the Tethys Sea closed, and the Neotethys basin appeared. Climates were warm, with no evidence of glaciation, as in the Triassic, there was apparently no land over either pole, and no extensive ice caps existed. In contrast, the North American Jurassic record is the poorest of the Mesozoic, the Jurassic was a time of calcite sea geochemistry in which low-magnesium calcite was the primary inorganic marine precipitate of calcium carbonate. Carbonate hardgrounds were thus very common, along with calcitic ooids, calcitic cements, the first of several massive batholiths were emplaced in the northern American cordillera beginning in the mid-Jurassic, marking the Nevadan orogeny. Important Jurassic exposures are found in Russia, India, South America, Japan, Australasia. As the Jurassic proceeded, larger and more groups of dinosaurs like sauropods and ornithopods proliferated in Africa. Middle Jurassic strata are well represented nor well studied in Africa. Late Jurassic strata are also poorly represented apart from the spectacular Tendaguru fauna in Tanzania, the Late Jurassic life of Tendaguru is very similar to that found in western North Americas Morrison Formation. During the Jurassic period, the primary living in the sea were fish
4.
Bird
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Birds, a subgroup of Reptiles, are the last living examples of Dinosaurs. Birds live worldwide and range in size from the 5 cm bee hummingbird to the 2.75 m ostrich. They rank as the class of tetrapods with the most living species, at ten thousand. Birds are the closest living relatives of crocodilians, the fossil record indicates that birds evolved from feathered ancestors within the theropod group of saurischian dinosaurs. True birds first appeared during the Cretaceous period, around 100 million years ago, Birds, especially those in the southern continents, survived this event and then migrated to other parts of the world while diversifying during periods of global cooling. Primitive bird-like dinosaurs that lie outside class Aves proper, in the broader group Avialae, have been found dating back to the mid-Jurassic period, around 170 million years ago. Birds have wings which are more or less developed depending on the species, the digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have evolved for swimming. Many species annually migrate great distances, Birds are social, communicating with visual signals, calls, and bird songs, and participating in such social behaviours as cooperative breeding and hunting, flocking, and mobbing of predators. The vast majority of species are socially monogamous, usually for one breeding season at a time, sometimes for years. Other species have breeding systems that are polygynous or, rarely, Birds produce offspring by laying eggs which are fertilised through sexual reproduction. They are usually laid in a nest and incubated by the parents, most birds have an extended period of parental care after hatching. Some birds, such as hens, lay eggs even when not fertilised, songbirds, parrots, and other species are popular as pets. Guano is harvested for use as a fertiliser, Birds prominently figure throughout human culture. About 120–130 species have become extinct due to human activity since the 17th century, human activity threatens about 1,200 bird species with extinction, though efforts are underway to protect them. Recreational birdwatching is an important part of the ecotourism industry, the first classification of birds was developed by Francis Willughby and John Ray in their 1676 volume Ornithologiae. Carl Linnaeus modified that work in 1758 to devise the taxonomic classification system currently in use, Birds are categorised as the biological class Aves in Linnaean taxonomy. Phylogenetic taxonomy places Aves in the dinosaur clade Theropoda, Aves and a sister group, the clade Crocodilia, contain the only living representatives of the reptile clade Archosauria
5.
Clade
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A clade is a group of organisms that consists of a common ancestor and all its lineal descendants, and represents a single branch on the tree of life. The common ancestor may be an individual, a population, a species, clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently, over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades, the term clade was coined in 1957 by the biologist Julian Huxley to refer to the result of cladogenesis, a concept Huxley borrowed from Bernhard Rensch. Many commonly named groups are clades, for example, rodents or insects, because in each case, their name refers to a common ancestor with all its descendant branches. Rodents, for example, are a branch of mammals that split off after the end of the period when the clade Dinosauria stopped being the dominant terrestrial vertebrates 66 million years ago, the original population and all its descendants are a clade. The rodent clade corresponds to the order Rodentia, and insects to the class Insecta and these clades include smaller clades, such as chipmunk or ant, each of which consists of even smaller clades. The clade rodent is in included in the mammal, vertebrate. The phenomenon of convergent evolution is, however, responsible for cases where there are misleading similarities in the morphology of groups that evolved from different lineages. The publication of Darwins theory of evolution in 1859 gave this view increasing weight, thomas Henry Huxley, an early advocate of evolutionary theory, proposed a revised taxonomy based on clades. For example, he grouped birds with reptiles, based on fossil evidence, german biologist Emil Hans Willi Hennig is considered to be the founder of cladistics. Taxonomists have increasingly worked to make the taxonomic system reflect evolution, when it comes to naming, however, this principle is not always compatible with the traditional rank-based nomenclature. For these and other reasons, phylogenetic nomenclature has been developed, a clade is by definition monophyletic, meaning that it contains one ancestor and all its descendants. The ancestor can be known or unknown, any and all members of a clade can be extant or extinct, the science that tries to reconstruct phylogenetic trees and thus discover clades is called phylogenetics or cladistics, the latter term coined by Ernst Mayr, derived from clade. The results of analyses are tree-shaped diagrams called cladograms, they. Three methods of defining clades are featured in phylogenetic nomenclature, node-, stem-, the relationship between clades can be described in several ways, A clade located within a clade is said to be nested within that clade. In the diagram, the clade, i. e. the apes. Two clades are sisters if they have a common ancestor
6.
Theropoda
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Theropoda or theropods are a group of saurischian dinosaurs. Theropods were ancestrally carnivorous, although a number of theropod groups evolved herbivory, omnivory, piscivory, in the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by 10,000 living species. Theropods exhibit a range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always considered the ancestral diet for theropods as a group. However, discoveries in the late 20th and early 21st centuries showed that a variety of diets existed even in more basal lineages, all early finds of theropod fossils showed them to be primarily carnivorous. For example, a Compsognathus longipes fossil was found with a lizard in its stomach, the first confirmed non-carnivorous fossil theropods found were the therizinosaurs, originally known as segnosaurs. First thought to be prosauropods, these enigmatic dinosaurs were later proven to be highly specialized, therizinosaurs possessed large abdomens for processing plant food, and small heads with beaks and leaf-shaped teeth. Further study of maniraptoran theropods and their relationships showed that therizinosaurs were not the early members of this group to abandon carnivory. Several other lineages of early maniraptors show adaptations for an omnivorous diet, oviraptorosaurs, ornithomimosaurs and advanced troodontids were likely omnivorous as well, and some early theropods appear to have specialized in catching fish. Diet is largely deduced by the morphology, tooth marks on bones of the prey. Some theropods, such as Baryonyx, Lourinhanosaurus, ornithomimosaurs, and birds, are known to use gastroliths, the majority of theropod teeth are blade-like, with serration on the edges, called ziphodont. Others are pachydont or phyllodont depending on the shape of the tooth or denticles, the morphology of the teeth is distinct enough to tell the major families apart, which indicate different diet strategies. The folds helped the teeth stay in place longer, especially as theropods evolved into larger sizes and had more force in their bite, Mesozoic theropods were also very diverse in terms of skin texture and covering. Feathers or feather-like structures are attested in most lineages of theropods, however, outside the coelurosaurs, feathers may have been confined to the young, smaller species, or limited parts of the animal. Many larger theropods had skin covered in small, bumpy scales, in some species, these were interspersed with larger scales with bony cores, or osteoderms. This type of skin is best known in the ceratosaur Carnotaurus, the coelurosaur lineages most distant from birds had feathers that were relatively short and composed of simple, possibly branching filaments. Simple filaments are also seen in therizinosaurs, which also possessed large, more fully feathered theropods, such as dromaeosaurs, usually retain scales only on the feet. Some species may have mixed feathers elsewhere on the body as well, scansoriopteryx preserved scales near the underside of the tail, and Juravenator may have been predominantly scaly with some simple filaments interspersed
7.
Dinosaur
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Dinosaurs are a diverse group of reptiles of the clade Dinosauria that first appeared during the Triassic. Although the exact origin and timing of the evolution of dinosaurs is the subject of active research and they became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201 million years ago. Their dominance continued through the Jurassic and Cretaceous periods and ended when the Cretaceous–Paleogene extinction event led to the extinction of most dinosaur groups 66 million years ago, until the late 20th century, all groups of dinosaurs were believed to be extinct. As such, birds were the dinosaur lineage to survive the mass extinction event. This article deals primarily with non-avian dinosaurs, Dinosaurs are a varied group of animals from taxonomic, morphological and ecological standpoints. Birds, at over 10,000 living species, are the most diverse group of vertebrates besides perciform fish, using fossil evidence, paleontologists have identified over 500 distinct genera and more than 1,000 different species of non-avian dinosaurs. Dinosaurs are represented on every continent by both extant species and fossil remains, while dinosaurs were ancestrally bipedal, many extinct groups included quadrupedal species, and some were able to shift between these stances. Elaborate display structures such as horns or crests are common to all dinosaur groups, evidence suggests that egg laying and nest building are additional traits shared by all dinosaurs.7 meters and heights of 18 meters and were the largest land animals of all time. Still, the idea that dinosaurs were uniformly gigantic is a misconception based in part on preservation bias, as large. Many dinosaurs were small, Xixianykus, for example, was only about 50 cm long. Through the first half of the 20th century, before birds were recognized to be dinosaurs, most of the community believed dinosaurs to have been sluggish. Most research conducted since the 1970s, however, has indicated that all dinosaurs were active animals with elevated metabolisms and numerous adaptations for social interaction. The large sizes of some groups, as well as their seemingly monstrous and fantastic nature, have ensured dinosaurs regular appearance in best-selling books and films. Persistent public enthusiasm for the animals has resulted in significant funding for dinosaur science, the term is derived from the Greek words δεινός and σαῦρος. Though the taxonomic name has often interpreted as a reference to dinosaurs teeth, claws. Instead, dinosaurs, like many forms of reptile sub-groups, did not exhibit characteristics which were traditionally regarded as reptilian. Under phylogenetic nomenclature, dinosaurs are usually defined as the group consisting of Triceratops, Neornithes, their most recent common ancestor, Birds are now recognized as being the sole surviving lineage of theropod dinosaurs. In traditional taxonomy, birds were considered a class that had evolved from dinosaurs
8.
Paraves
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Paraves are a widespread group of theropod dinosaurs that originated in the Late Jurassic period. In addition to the extinct dromaeosaurids, troodontids, and scansoriopterygids, primitive members of Paraves are well-known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. Like other theropods, all paravians are bipedal, walking on their two hind legs, the teeth of primitive paravians were curved and serrated, but not blade-like except in some specialized species such as Dromaeosaurus albertensis. The serrations on the front edge of dromaeosaurid and troodontid teeth were small and fine, while the back edge had serrations which were very large. Paravians generally have long, winged forelimbs, though these have become smaller in many flightless species, the wings usually bore three large, flexible, clawed fingers in early forms. The fingers became fused and stiffened and the highly reduced or lost in some advanced lineages. Many early members of Paraves had both well-developed wings and long feathers on the legs, which in some cases, formed a second set of airfoils. These species, most famously represented by Microraptor gui, have often referred to as four winged dinosaurs. Microraptor in particular seems to represent a case of flight evolving independently of the bird lineage within Paraves. The enlarged second toe bore a large, curved sickle-shaped claw. This claw was especially large and flattened from side to side in the large-bodied predatory eudromaeosaurs, in these early species, the first toe was usually small and angled inward toward the center of the body, but only became fully reversed in more specialized members of the bird lineage. One species, Balaur bondoc, possessed a first toe which was modified in parallel with the second. Both the first and second toes on each foot of B. bondoc were held retracted, initial speculation regarded the claws as slashing implements used to disembowel large prey. In this scenario, the upper foot would serve as an anchor point for powerful tendons to improve kicking ability. However, subsequent studies of the actual claw shape showed that the underside of the claw was only weakly keeled, instead, it appeared to be more of a hooking implement. Manning et al. suggested in 2006 that the claws were similar to crampons and were used for climbing and they compared the claw and overall foot anatomy of various primitive species with modern birds to shed light on their actual function. Fowler and colleagues showed that many modern birds also have enlarged claws on the second toes. Other lines of evidence for this behavior include teeth which had large, in more primitive dromaeosaurids and in troodontids, the feet were not as specialized and the claws were not as large or as hooked
9.
Class (biology)
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In biological classification, class is, a taxonomic rank. Other well-known ranks in descending order of size are life, domain, kingdom, phylum, order, family, genus, as for the other well-known ranks, there is the option of an immediately lower rank, indicated by the prefix sub-, subclass. A taxonomic unit, a taxon, in that rank, in that case the plural is classes The composition of each class is determined by a taxonomist. Often there is no agreement, with different taxonomists taking different positions. There are no rules that a taxonomist needs to follow in describing a class. For example, dogs are assigned to the phylum Chordata, in the class Mammalia. In botany, classes are now rarely discussed, for some clades, a number of alternative classifications are used. In the first edition of his Systema Naturae, carl Linnaeus divided all three of his kingdoms of Nature into classes. The class was considered the highest level of the hierarchy until George Cuviers embranchements. Cladistics List of animal classes Phylogenetics Systematics Taxonomy
10.
Order (biology)
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In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, an immediately higher rank, superorder, may be added directly above order, while suborder would be a lower rank. A taxonomic unit, a taxon, in that rank, in that case the plural is orders. Example, The Juglans and Hickory trees belong to the family Juglandaceae, what does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no agreement, with different taxonomists each taking a different position. There are no rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognised only rarely, for some groups of organisms, consistent suffixes are used to denote that the rank is an order. The Latin suffix -formes meaning having the form of is used for the name of orders of birds and fishes. The suffix -ales is for the name of orders of plants, fungi, for some clades covered by the International Code of Zoological Nomenclature, a number of additional classifications are sometimes used, although not all of these are officially recognised. In their 1997 classification of mammals, McKenna and Bell used two levels between superorder and order, grandorder and mirorder. Michael Novacek inserted them at the same position, michael Benton inserted them between superorder and magnorder instead. This position was adopted by Systema Naturae 2000 and others, in botany, the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined, the superorder rank is commonly used, with the ending -anae that was initiated by Armen Takhtajans publications from 1966 onwards. Carl Linnaeus was the first to apply it consistently to the division of all three kingdoms of nature in his Systema Naturae, for plants, Linnaeus orders in the Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. In French botanical publications, from Michel Adansons Familles naturelles des plantes and until the end of the 19th century, some of the plant families still retain the names of Linnaean natural orders or even the names of pre-Linnaean natural groups recognised by Linnaeus as orders in his natural classification. Such names are known as family names. In zoology, the Linnaean orders were used more consistently and that is, the orders in the zoology part of the Systema Naturae refer to natural groups. Some of his names are still in use
11.
Crocodilia
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The Crocodilia is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 83.5 million years ago in the Late Cretaceous period and are the closest living relatives of birds, as the two groups are the only known survivors of the Archosauria. Members of the total group, the clade Pseudosuchia, appeared about 250 million years ago in the Early Triassic period. The order Crocodilia includes the true crocodiles, the alligators and caimans, although the term crocodiles is sometimes used to refer to all of these, crocodilians is a less ambiguous vernacular term for members of this group. Large, solidly built, lizard-like reptiles, crocodilians have flattened snouts, laterally compressed tails, and eyes, ears. They swim well and can move on land in a high walk and their skin is thick and covered in non-overlapping scales. They have conical, peg-like teeth and a powerful bite and they have a four-chambered heart and, somewhat like birds, a unidirectional looping system of airflow within the lungs, but like other non-avian reptiles they are ectotherms. Crocodilians are found mainly in lowlands in the tropics, but alligators also live in the southeastern United States, Crocodilians are typically solitary and territorial, though cooperative feeding does occur. During breeding, dominant males try to monopolise available females, females lay eggs in holes or in mounds and, unlike most other reptiles, care for their hatched young. Eight species of crocodilians are known to have attacked humans, the largest number of attacks comes from the Nile crocodile. Artistic and literary representations of crocodilians have appeared in cultures around the world since at least Ancient Egypt. Crocodilia and Crocodylia have been used interchangeably for decades starting with Schmidts redescription of the group from the defunct term Loricata. Schmidt used the older term Crocodilia, based on Owens original name for the group, shortly after, Wermuth opted for Crocodylia as the proper name for this redescribed group, basing it on the type genus Crocodylus. Dundee—in a revision of many reptilian and amphibian names—argued strongly for Crocodylia to be the spelling for the group, however, it was not until the advent of cladistics and phylogenetic nomenclature that a more solid justification for assuming one spelling over the other was proposed. Prior to 2003, Crocodilia/Crocodylia was a group that encompassed the modern-day animals as well as their distant relatives now in the larger groups called Crocodylomorpha. Under its current definition Crocodylia is restricted to only the most recent ancestor of todays modern-day crocodilians and this distinction is more important for paleontologists studying crocodilian evolution. As such, the alternate spellings Crocodilia and Crocodylia are still used interchangeably in the neontological literature, Crocodilia appears to be a Latinizing of the Greek κροκόδειλος, which means both lizard and Nile crocodile. Crocodylia, as coined by Wermuth, in regards to the genus Crocodylus appears to be derived from the ancient Greek κρόκη —meaning shingle or pebble—and δρîλος or δρεîλος for worm, the name may refer to the animals habit of basking on the pebbled shores of the Nile
12.
Reptile
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Reptiles are tetrapod animals in the class Reptilia, comprising todays turtles, crocodilians, snakes, amphisbaenians, lizards, tuatara, and their extinct relatives. The study of these traditional reptile orders, historically combined with that of amphibians, is called herpetology. Because some reptiles are more related to birds than they are to other reptiles. For this reason, many scientists prefer to consider the birds part of Reptilia as well. Some early examples include the lizard-like Hylonomus and Casineria, in addition to the living reptiles, there are many diverse groups that are now extinct, in some cases due to mass extinction events. In particular, the K–Pg extinction wiped out the pterosaurs, plesiosaurs, ornithischians, and sauropods, as well as species of theropods, crocodyliforms. Modern non-avian reptiles inhabit every continent with the exception of Antarctica, several living subgroups are recognized, Testudines, approximately 400 species, Sphenodontia,1 species, Squamata, over 9,600 species, Crocodilia,25 species, and Aves,10,000 species. Reptiles are tetrapod vertebrates, creatures that either have four limbs or, unlike amphibians, reptiles do not have an aquatic larval stage. As amniotes, reptile eggs are surrounded by membranes for protection and transport, many of the viviparous species feed their fetuses through various forms of placenta analogous to those of mammals, with some providing initial care for their hatchlings. In the 18th century, the reptiles were, from the outset of classification, the terms reptile and amphibian were largely interchangeable, reptile being preferred by the French. Josephus Nicolaus Laurenti was the first to use the term Reptilia for an expanded selection of reptiles. Today, the two groups are commonly treated under the same heading as herptiles. He subsequently proposed the names of Sauropsida and Ichthyopsida for the two groups. In 1866, Haeckel demonstrated that vertebrates could be divided based on their strategies, and that reptiles, birds. The terms Sauropsida and Theropsida were used again in 1916 by E. S, Goodrich to distinguish between lizards, birds, and their relatives on the one hand and mammals and their extinct relatives on the other. Goodrich supported this division by the nature of the hearts and blood vessels in each group, according to Goodrich, both lineages evolved from an earlier stem group, Protosauria in which he included some animals today considered reptile-like amphibians, as well as early reptiles. Watson observed that the first two groups diverged very early in history, so he divided Goodrichs Protosauria between them. He also reinterpreted Sauropsida and Theropsida to exclude birds and mammals, thus his Sauropsida included Procolophonia, Eosuchia, Millerosauria, Chelonia, Squamata, Rhynchocephalia, Crocodilia, thecodonts, non-avian dinosaurs, pterosaurs, ichthyosaurs, and sauropterygians
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Archosaur
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Archosaurs are a group of diapsid amniotes whose living representatives consist of birds and crocodilians. This group also includes all extinct dinosaurs, extinct crocodilian relatives. Archosauria, the clade, is a crown group that includes the most recent common ancestor of living birds. It includes two clades, Pseudosuchia, which includes crocodilians and their extinct relatives, and Ornithosuchia. Archosaurs can be distinguished from other tetrapods on the basis of synapomorphies, or shared characteristics. The simplest and most widely agreed synapomorphies of archosaurs include teeth set in sockets, antorbital and mandibular fenestrae, being set in sockets, the teeth were less likely to be torn loose during feeding. This feature is responsible for the name thecodont, which used to apply to many Triassic archosaurs. Some archosaurs, such as birds, are secondarily toothless, antorbital fenestrae reduced the weight of the skull, which was relatively large in early archosaurs, rather like that of modern crocodilians. Mandibular fenestrae may also have reduced the weight of the jaw in some forms, the fourth trochanter provides a large site for the attachment of muscles on the femur. There is some debate about when archosaurs first appeared and those who classify the Permian reptiles Archosaurus rossicus and/or Protorosaurus speneri as true archosaurs maintain that archosaurs first appeared in the late Permian. The earliest archosaurs were rauisuchians, such as Scythosuchus and Tsylmosuchus, synapsids were the dominant land vertebrates throughout the Permian, but most perished in the Permian-Triassic extinction event. Very few large synapsids survived the event, although one form, Lystrosaurus, but archosaurs quickly became the dominant land vertebrates in the early Triassic. The two most commonly suggested explanations for this are, Archosaurs made more progress towards erect limbs than synapsids. An objection to this explanation is that became dominant while they still had sprawling or semi-erect limbs, similar to those of Lystrosaurus. The early Triassic was predominantly arid, because most of the land was concentrated in the supercontinent Pangaea. Archosaurs were probably better at conserving water than early synapsids because, Modern diapsids excrete uric acid and it is reasonable to suppose that archosaurs also excreted uric acid, and therefore were good at conserving water. The aglandular skins of diapsids would also have helped to conserve water, Modern mammals excrete urea, which requires a lot of water to keep it dissolved. Their skins also contain many glands, which also lose water, assuming that early synapsids had similar features, e. g. as argued by the authors of Palaeos, they were at a disadvantage in a mainly arid world
14.
Palaeognathae
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Palaeognathae, or paleognaths, is one of the two living clades of birds – the other being Neognathae. Together, these two form the clade Neornithes. Palaeognathae contains five extant branches of flightless lineages, termed ratites, and one flying lineage, there are 47 species of tinamous,5 of kiwis,3 of cassowaries,1 of emus,2 of rheas and 2 of ostrich. There are three groups, the Lithornithiformes, the Dinornithiformes and the Aepyornithiformes, that are undisputed members of Palaeognathae. There are other extinct birds which have been allied with the Palaeognathae by at least one author, but their affinities are a matter of dispute. The word Paleognath is derived from the ancient Greek for old jaws in reference to the anatomy of the palate. There are also other scientific controversies about their evolution. Given the northern location of the morphologically most basal fossil forms. One study of molecular and paleontological data found that modern bird orders, including the paleognathous ones, benton summarized this and other molecular studies as implying that paleognaths should have arisen 110 to 120 million years ago in the Early Cretaceous. He points out, however, that there is no fossil record until 70 million years ago, unrelated birds might have developed ratite-like anatomies multiple times around the world through convergent evolution. McDowell asserted that the similarities in the anatomy of paleognathes might actually be neoteny. He noted that there were other feature of the skull, such as the retention of sutures into adulthood, thus, perhaps the characteristic palate was actually a frozen stage that many carinate bird embryos passed through during development. The retention of early stages, then, may have been a mechanism by which various birds became flightless. Hope reviewed all known fossils from the Mesozoic looking for evidence of the origin of the evolutionary radiation of the Neornithes. That radiation would also signal that the paleognaths had already diverged and she notes five Early Cretaceous taxa that have been assigned to the Palaeognathae. She finds that none of them can be assigned as such. However, she find evidence that the Neognathae and, therefore. Vegavis is a bird from the Maastrichtian period of Late Cretaceous Antarctica
15.
Anseriformes
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In fact, these living species are all included in Anatidae except for the three screamers and the magpie goose. All species in the order are highly adapted for an aquatic existence at the water surface, the males, except for the screamers, also have a penis, a trait that has disappeared in Neoaves. All are web-footed for efficient swimming, the earliest known Anseriform is the recently discovered Vegavis, which lived during the Cretaceous period. It is thought that the Anseriformes originated when the original Galloanserae split into the two main lineages, the extinct dromornithids may possibly represent early offshoots of the anseriform line, if they arent stem-Galliformes instead, and so maybe Gastornis. The ancestors of the Anseriformes developed the characteristic bill structure that they still share, the prehistoric wading presbyornithids were even more bizarre. The Anseriformes and the Galliformes are the most primitive neognathous birds, Anatidae systematics, especially regarding placement of some odd genera in the dabbling ducks or shelducks, is not fully resolved. See the Anatidae article for information, and for alternate taxonomic approaches. Some unusual groups, such as the extinct Gastornithidae and Dromornithidae, are found to be at the base of the Anseriformes family tree. Anatidae is traditionally divided into subfamilies Anatinae and Anserinae, the Anatinae consists of tribes Anatini, Aythyini, Mergini and Tadornini. The higher-order classification below follows a phylogenetic analysis performed by Angolin,2007, Mikkos Phylogeny Archive, the clade Somaterini has a single genus Somateria. List of Anseriformes by population Agnolin, F. Brontornis burmeisteri Moreno & Mercerat, un Anseriformes gigante del Mioceno Medio de Patagonia, revista del Museo Argentino de Ciencias Naturales. Clarke, J. A. Tambussi, C. P. Noriega, erickson, G. M. & Ketcham, R. A. Definitive fossil evidence for the extant avian radiation in the Cretaceous, & Zusi, R. L. Higher-order phylogeny of modern birds based on comparative anatomy. Zoological Journal of the Linnen Society, murray, P. F. & Vickers-Rich, P. Magnificent Mihirungs, The Colossal Flightless Birds of the Australian Dreamtime
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Galliformes
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The name derives from gallus, Latin for cock or rooster. Common names are gamefowl or gamebirds, landfowl, gallinaceous birds, wildfowl or just fowl are also often used for the Galliformes, but usually these terms also refer to waterfowl, and occasionally to other commonly hunted birds. This group has about 290 species, one or more of which are found in every part of the worlds continents. They are rarer on islands, and in contrast to the closely related waterfowl, are absent from oceanic islands—unless introduced there by humans. Several species have been domesticated during their long and extensive relationships with humans and this order contains five families, Phasianidae, Odontophoridae, Numididae, Cracidae, and Megapodiidae. They are important as seed dispersers and predators in the ecosystems they inhabit, many gallinaceous species are skilled runners and escape predators by running rather than flying. Males of most species are more colorful than the females, males often have elaborate courtship behaviors that include strutting, fluffing of tail or head feathers, and vocal sounds. The living Galliformes were once divided into seven or more families, despite their distinctive appearance, grouse and turkeys probably do not warrant separation as families due to their recent origin from partridge- or pheasant-like birds. The turkeys became larger after their ancestors colonized temperate and subtropical North America, the ancestors of grouse, though, adapted to harsh climates and could thereby colonize subarctic regions. Consequently, the Phasianidae are expanded in current taxonomy to include the former Tetraonidae and Meleagrididae as subfamilies, the Anseriformes and the Galliformes together make up the Galloanserae. They are basal among the living birds, and normally follow the Paleognathae in modern bird classification systems. This was first proposed in the Sibley-Ahlquist taxonomy and has been the one major change of that proposed scheme that was almost universally adopted. However, the Galliformes as they were traditionally delimited are called Gallomorphae in the Sibley-Ahlquist taxonomy and this is not a natural group, however, but rather an erroneous result of the now-obsolete phenetic methodology employed in the Sibley-Ahlquist taxonomy. Phenetic studies do not distinguish between plesiomorphic and apomorphic characters, which leads to basal lineages appearing as monophyletic groups, historically, the buttonquails, mesites and the hoatzin were placed in the Galliformes, too. The former are now known to be adapted to an inland lifestyle, whereas the mesites are probably closely related to pigeons. The relationships of the hoatzin are entirely obscure, and it is treated as a monotypic order Opisthocomiformes to signify this. The earliest galliform-like fossils hail from the Late Cretaceous, most notably those of Austinornis lentus and its partial left tarsometatarsus was found in the Austin Chalk near Fort McKinney, Texas, dating to about 85 million years ago. This bird was quite closely related to Galliformes, but whether it was a part of these or belongs elsewhere in the little-known galliform branch of Galloanserae is not clear
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Neoaves
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Neoaves is a clade that consists of all modern birds with the exception of Paleognathae and Galloanserae. The early diversification of the various neoavian groups occurred very rapidly, one hypothesis for the phylogeny of modern birds was presented by Prum, R. O. et al. The following cladogram illustrates the relationships, with some taxon names following Yury
18.
Phylogenetics
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In biology, phylogenetics /ˌfaɪloʊdʒəˈnɛtɪks, -lə-/ is the study of the evolutionary history and relationships among individuals or groups of organisms. These relationships are discovered through phylogenetic inference methods that evaluate observed heritable traits, the result of these analyses is a phylogeny – a diagrammatic hypothesis about the history of the evolutionary relationships of a group of organisms. The tips of a tree can be living organisms or fossils. Phylogenetic analyses have become central to understanding biodiversity, evolution, ecology, taxonomy is the classification, identification and naming of organisms. It is usually richly informed by phylogenetics, but remains a methodologically and logically distinct discipline, usual methods of phylogenetic inference involve computational approaches implementing the optimality criteria and methods of parsimony, maximum likelihood, and MCMC-based Bayesian inference. All these depend upon an implicit or explicit mathematical model describing the evolution of characters observed, prior to 1990, phylogenetic inferences were generally presented as narrative scenarios. Such methods are often ambiguous and lack explicit criteria for evaluating alternative hypotheses, the term phylogeny derives from the German Phylogenie, introduced by Haeckel in 1866, and the Darwinian approach to classification became known as the phyletic approach. During the late 19th century, Ernst Haeckels recapitulation theory, or biogenetic fundamental law, was widely accepted, but this theory has long been rejected. O. See, Huerta-Cepas, Jaime, Dopazo, Joaquín, Gabaldón, ETE, A python Environment for Tree Exploration. PhylomeDB, A public database hosting thousands of gene phylogenies ranging many different species, See, Huerta-Cepas, J. Capella-Gutierrez, S. Pryszcz, L. P. Denisov, I. Kormes, D. Marcet-Houben, M. Gabaldón, T. PhylomeDB v3.0, An expanding repository of genome-wide collections of trees, alignments, lents, N. H. Cifuentes, O. E. Carpi, A. Teaching the Process of Molecular Phylogeny and Systematics, A Multi-Part Inquiry-Based Exercise
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House sparrow
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The house sparrow is a bird of the sparrow family Passeridae, found in most parts of the world. A small bird, it has a length of 16 cm. Females and young birds are coloured brown and grey, and males have brighter black, white. One of about 25 species in the genus Passer, the sparrow is native to most of Europe, the Mediterranean region. Its intentional or accidental introductions to many regions, including parts of Australia, Africa, the house sparrow is strongly associated with human habitations, and can live in urban or rural settings. Though found in varied habitats and climates, it typically avoids extensive woodlands, grasslands. It feeds mostly on the seeds of grains and weeds, but it is an opportunistic eater and commonly eats insects and its predators include domestic cats, hawks, owls, and many other predatory birds and mammals. Because of its numbers, ubiquity, and association with human settlements, though it is widespread and abundant, its numbers have declined in some areas. The animals conservation status is listed as least concern on the IUCN Red List, the house sparrow is typically about 16 cm long, ranging from 14 to 18 cm. It is a bird with a full chest and a large. Its bill is stout and conical with a length of 1. 1–1.5 cm. Its tail is short, at 5. 2–6.5 cm long, the wing chord is 6. 7–8.9 cm, and the tarsus is 1. 6–2.5 cm. In mass, the house sparrow ranges from 24 to 39.5 g, females usually are slightly smaller than males. The median mass on the European continent for both sexes is about 30 g, and in more southerly subspecies is around 26 g, younger birds are smaller, males are larger during the winter, and females are larger during the breeding season. Birds at higher latitudes, colder climates, and sometimes higher altitudes are larger, the plumage of the house sparrow is mostly different shades of grey and brown. The sexes exhibit strong dimorphism, the female is mostly buffish above and below, while the male has boldly coloured head markings, a reddish back, and grey underparts. The male has a grey crown from the top of its bill to its back. It has black around its bill, on its throat, and it has a small white stripe between the lores and crown and small white spots immediately behind the eyes, with black patches below and above them
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Timeline of the evolutionary history of life
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This timeline of the evolutionary history of life represents the current scientific theory outlining the major events during the development of life on planet Earth. In biology, evolution is any change across generations in the heritable characteristics of biological populations. Evolutionary processes give rise to diversity at every level of organization, from kingdoms to species. The similarities between all present day organisms indicate the presence of an ancestor from which all known species, living. More than 99 percent of all species, amounting to five billion species. Estimates on the number of Earths current species range from 10 million to 14 million, of which about 1.2 million have been documented and over 86 percent have not yet been described. Species go extinct constantly as environments change, organisms compete for environmental niches, occasionally biodiversity on the planet takes a hit in the form of a mass extinction in which the extinction rate is much higher than usual. A large extinction event often represents an accumulation of smaller extinction events that take place in a brief period of time. The first known mass extinction in earths history was the Great Oxygenation Event 2.4 billion years ago, the event led to the loss of most of the planets obligate anaerobes. The Holocene extinction event is currently under way, in this timeline, Ma means million years ago, ka means thousand years ago, and ya means years ago. 4000 Ma –2500 Ma 2500 Ma –542 Ma, contains the Palaeoproterozoic, Mesoproterozoic and Neoproterozoic eras. 542 Ma – present The Phanerozoic Eon, literally the period of well-displayed life, marks the appearance in the record of abundant. It is subdivided into three eras, the Paleozoic, Mesozoic and Cenozoic, which are divided by mass extinctions. 542 Ma –251.0 Ma and contains the Cambrian, Ordovician, Silurian, Devonian, Carboniferous, from 251.4 Ma to 66 Ma and containing the Triassic, Jurassic and Cretaceous periods. 66 Ma – present Dawkins, Richard, the Ancestors Tale, A Pilgrimage to the Dawn of Life. Understanding Evolution, your one-stop resource for information on evolution, explore complete phylogenetic tree interactively Brandt, Niel. Houston, TX, The TalkOrigins Foundation, Inc, interactive timeline from Big Bang to present Plant Evolution. Sequence of Plant Evolution The History of Animal Evolution, sequence of Animal Evolution Yeo, Dannel, Drage, Thomas
21.
Water
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Water is a transparent and nearly colorless chemical substance that is the main constituent of Earths streams, lakes, and oceans, and the fluids of most living organisms. Its chemical formula is H2O, meaning that its molecule contains one oxygen, Water strictly refers to the liquid state of that substance, that prevails at standard ambient temperature and pressure, but it often refers also to its solid state or its gaseous state. It also occurs in nature as snow, glaciers, ice packs and icebergs, clouds, fog, dew, aquifers, Water covers 71% of the Earths surface. It is vital for all forms of life. Only 2. 5% of this water is freshwater, and 98. 8% of that water is in ice and groundwater. Less than 0. 3% of all freshwater is in rivers, lakes, and the atmosphere, a greater quantity of water is found in the earths interior. Water on Earth moves continually through the cycle of evaporation and transpiration, condensation, precipitation. Evaporation and transpiration contribute to the precipitation over land, large amounts of water are also chemically combined or adsorbed in hydrated minerals. Safe drinking water is essential to humans and other even though it provides no calories or organic nutrients. There is a correlation between access to safe water and gross domestic product per capita. However, some observers have estimated that by 2025 more than half of the population will be facing water-based vulnerability. A report, issued in November 2009, suggests that by 2030, in developing regions of the world. Water plays an important role in the world economy, approximately 70% of the freshwater used by humans goes to agriculture. Fishing in salt and fresh water bodies is a source of food for many parts of the world. Much of long-distance trade of commodities and manufactured products is transported by boats through seas, rivers, lakes, large quantities of water, ice, and steam are used for cooling and heating, in industry and homes. Water is an excellent solvent for a variety of chemical substances, as such it is widely used in industrial processes. Water is also central to many sports and other forms of entertainment, such as swimming, pleasure boating, boat racing, surfing, sport fishing, Water is a liquid at the temperatures and pressures that are most adequate for life. Specifically, at atmospheric pressure of 1 bar, water is a liquid between the temperatures of 273.15 K and 373.15 K
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Unicellular organism
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A unicellular organism, also known as a single-celled organism, is an organism that consists of only one cell, unlike a multicellular organism that consists of more than one cell. Historically, simple unicellular organisms have been referred to as monads, though this term is used more specifically to describe organisms of the genus Monas. The main groups of organisms are bacteria, archaea, protozoa, unicellular algae. Unicellular organisms fall into two categories, prokaryotic organisms and eukaryotic organisms. Unicellular organisms are thought to be the oldest form of life, prokaryotes, most Protista, and some fungi are unicellular. Although some of these live in colonies, they dont exhibit specialization. These organisms live together, and each cell in the colony is the same, however, each individual cell must carry out all life processes to survive. In contrast, even the simplest multicellular organisms have cells that depend on each other to survive, most multicellular organisms have a unicellular life-cycle stage. Gametes, for example, are reproductive unicells for multicellular organisms, additionally, multicellularity appears to have evolved independently many times in the history of life. Some organisms are partially uni- and multicellular, like Dictyostelium discoideum, additionally, unicellular organisms can be multinucleate, like Myxogastria and Plasmodium. Candidatus Magnetoglobus multicellular, related to Deltaproteobacteria, is a multicellular prokaryote and it is neither unicellular, nor a colony. Primitive cells, often referred to as protocells, are the precursors to todays unicellular organisms, the RNA world hypothesis assumes that RNA molecules could form in abiotic conditions, which would require nucleic acids and ribose to be present. Theoretical and experimental findings show that nucleic acids and sugars could have been synthesized in early prebiotic conditions, compartmentalization was necessary for chemical reactions to be more likely as well as to differentiate reactions with the external environment. For example, an early RNA replicator ribozyme may have replicated other replicator ribozymes of different RNA sequences if not kept separate, when amphiphiles like lipids are placed in water, the hydrophobic tails aggregate to form micelles and vesicles, with the hydrophilic ends facing outwards. Primitive cells likely used self-assembling fatty-acid vesicles to separate chemical reactions, because of their simplicity and ability to self-assemble in water, its likely that these simple membranes predated other forms of early biological molecules. Prokaryotes lack membrane-bound organelles, such as mitochondria or a nucleus, instead, most prokaryotes have an irregular region that contains DNA, known as the nucleoid. Most prokaryotes have a single, circular chromosome, which is in contrast to eukaryotes, nutritionally, prokaryotes have the ability to utilize a wide range of organic and inorganic material for use in metabolism, including sulfur, cellulose, ammonia, or nitrite. Prokaryotes as a whole are ubiquitous in the environment and exist in extreme environments as well, bacteria are one of the world’s oldest forms of life, and are found virtually everywhere in nature
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Evolution of photosynthesis
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The first photosynthetic organisms probably evolved early in the evolutionary history of life and most likely used reducing agents such as hydrogen or electrons, rather than water. There are three major metabolic pathways by which photosynthesis is carried out, C3 photosynthesis, C4 photosynthesis, C3 photosynthesis is the oldest and most common form. The biochemical capacity to use water as the source for electrons in photosynthesis evolved in an ancestor of extant cyanobacteria. The geological record indicates that this event took place early in Earths history, at least 2450–2320 million years ago. Available evidence from studies of Archean sedimentary rocks indicates that life existed 3500 Ma. A clear paleontological window on cyanobacterial evolution opened about 2000 Ma, cyanobacteria remained principal primary producers throughout the Proterozoic Eon, in part because the redox structure of the oceans favored photoautotrophs capable of nitrogen fixation. Cyanobacteria remain critical to marine ecosystems as primary producers in oceanic gyres, as agents of biological nitrogen fixation, and, in modified form, as the plastids of marine algae. Early photosynthetic systems, such as those from green and purple sulfur and green and purple bacteria, are thought to have been anoxygenic. Green and purple sulfur bacteria are thought to have used hydrogen, green nonsulfur bacteria used various amino and other organic acids. Purple nonsulfur bacteria used a variety of organic molecules. Fossils of what are thought to be filamentous photosynthetic organisms have been dated at 3.4 billion years old, the main source of oxygen in the atmosphere is oxygenic photosynthesis, and its first appearance is sometimes referred to as the oxygen catastrophe. Geological evidence suggests that oxygenic photosynthesis, such as that in cyanobacteria, modern photosynthesis in plants and most photosynthetic prokaryotes is oxygenic. Oxygenic photosynthesis uses water as a donor, which is oxidized to molecular oxygen in the photosynthetic reaction center. Several groups of animals have formed relationships with photosynthetic algae. These are most common in corals, sponges and sea anemones and it is presumed that this is due to the particularly simple body plans and large surface areas of these animals compared to their volumes. This allows the mollusks to survive solely by photosynthesis for several months at a time, some of the genes from the plant cell nucleus have even been transferred to the slugs, so that the chloroplasts can be supplied with proteins that they need to survive. An even closer form of symbiosis may explain the origin of chloroplasts, chloroplasts have many similarities with photosynthetic bacteria, including a circular chromosome, prokaryotic-type ribosomes, and similar proteins in the photosynthetic reaction center. The endosymbiotic theory suggests that bacteria were acquired by early eukaryotic cells to form the first plant cells
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Eukaryote
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A eukaryote is any organism whose cells contain a nucleus and other organelles enclosed within membranes. Eukaryotes belong to the taxon Eukarya or Eukaryota, the presence of a nucleus gives eukaryotes their name, which comes from the Greek εὖ and κάρυον. Eukaryotic cells also contain other membrane-bound organelles such as mitochondria and the Golgi apparatus, in addition, plants and algae contain chloroplasts. Eukaryotic organisms may be unicellular or multicellular, only eukaryotes form multicellular organisms consisting of many kinds of tissue made up of different cell types. Eukaryotes can reproduce asexually through mitosis and sexually through meiosis and gamete fusion. In mitosis, one cell divides to produce two identical cells. In meiosis, DNA replication is followed by two rounds of division to produce four daughter cells each with half the number of chromosomes as the original parent cell. These act as sex cells resulting from genetic recombination during meiosis, the domain Eukaryota appears to be monophyletic, and so makes up one of the three domains of life. The two other domains, Bacteria and Archaea, are prokaryotes and have none of the above features, eukaryotes represent a tiny minority of all living things. However, due to their larger size, eukaryotes collective worldwide biomass is estimated at about equal to that of prokaryotes. Eukaryotes first developed approximately 1. 6–2.1 billion years ago, in 1905 and 1910, the Russian biologist Konstantin Mereschkowsky argued three things about the origin of nucleated cells. Firstly, plastids were reduced cyanobacteria in a symbiosis with a non-photosynthetic host, secondly, the host had earlier in evolution formed by symbiosis between an amoeba-like host and a bacteria-like cell that formed the nucleus. Thirdly, plants inherited photosynthesis from cyanobacteria, the split between the prokaryotes and eukaryotes was introduced in the 1960s. The concept of the eukaryote has been attributed to the French biologist Edouard Chatton, the terms prokaryote and eukaryote were more definitively reintroduced by the Canadian microbiologist Roger Stanier and the Dutch-American microbiologist C. B. van Niel in 1962. In his 1938 work Titres et Travaux Scientifiques, Chatton had proposed the two terms, calling the bacteria prokaryotes and organisms with nuclei in their cells eukaryotes. However he mentioned this in one paragraph, and the idea was effectively ignored until Chattons statement was rediscovered by Stanier. In 1967, Lynn Margulis provided microbiological evidence for endosymbiosis as the origin of chloroplasts and mitochondria in cells in her paper. In the 1970s, Carl Woese explored microbial phylogenetics, studying variations in 16S ribosomal RNA and this helped to uncover the origin of the eukaryotes and the symbiogenesis of two important eukaryote organelles, mitochondria and chloroplasts
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Multicellular organism
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Multicellular organisms are organisms that consist of more than one cell, in contrast to unicellular organisms. Multicellular organisms arise in different ways, for example by cell division or by aggregation of many single cells. Colonial organisms are the result of identical individuals joining together to form a colony. Multicellularity has evolved independently at least 46 times, including in some prokaryotes, like cyanobacteria, myxobacteria, actinomycetes, however, complex multicellular organisms evolved only in six eukaryotic groups, animals, fungi, brown algae, red algae, green algae, and land plants. It evolved repeatedly for Chloroplastida, once or twice for animals, once for brown algae, the first evidence of multicellularity is from cyanobacteria-like organisms that lived 3–3.5 billion years ago. To reproduce, true multicellular organisms must solve the problem of regenerating a whole organism from germ cells, animals have evolved a considerable diversity of cell types in a multicellular body, compared with 10–20 in plants and fungi. Loss of multicellularity occurred in some groups, fungi are predominantly multicellular, though early diverging lineages are largely unicellular and there have been numerous reversions to unicellularity across fungi. It may also have occurred in some red algae, but it is possible that they are primitively unicellular), loss of multicellularity is also considered probable in some green algae. In other groups, generally parasites, a reduction of multicellularity occurred, multicellular organisms, especially long-living animals, face the challenge of cancer, which occurs when cells fail to regulate their growth within the normal program of development. Changes in tissue morphology can be observed during this process, cancer in animals has often been described as a loss of multicellularity. There is a discussion about the possibility of existence of cancer in other organisms or even in protozoa. For example, plant galls have been characterized as tumors but some argue that plants do not develop cancer. In some multicellular groups, which are called Weismannists, a separation between a sterile somatic cell line and a cell line evolved. However, Weismannist development is relatively rare, as part of species have the capacity for somatic embryogenesis. One hypothesis for the origin of multicellularity is that a group of function-specific cells aggregated into a mass called a grex. This is essentially what slime molds do, another hypothesis is that a primitive cell underwent nucleus division, thereby becoming a coenocyte. A membrane would then form around each nucleus, thereby resulting in a group of connected cells in one organism and this is what plant and animal embryos do as well as colonial choanoflagellates. Because the first multicellular organisms were simple, soft organisms lacking bone, shell or other body parts
26.
Arthropod
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An arthropod is an invertebrate animal having an exoskeleton, a segmented body, and jointed appendages. Arthropods form the phylum Arthropoda, which includes the insects, arachnids, myriapods, arthropods are characterized by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages, the rigid cuticle inhibits growth, so arthropods replace it periodically by moulting. Their versatility has enabled them to become the most species-rich members of all guilds in most environments. They have over a million described species, making up more than 80% of all described living species, some of which. Arthropods range in size from the microscopic crustacean Stygotantulus up to the Japanese spider crab, arthropods primary internal cavity is a hemocoel, which accommodates their internal organs, and through which their haemolymph – analogue of blood – circulates, they have open circulatory systems. Like their exteriors, the organs of arthropods are generally built of repeated segments. Their nervous system is ladder-like, with paired ventral nerve cords running through all segments and their heads are formed by fusion of varying numbers of segments, and their brains are formed by fusion of the ganglia of these segments and encircle the esophagus. The respiratory and excretory systems of arthropods vary, depending as much on their environment as on the subphylum to which they belong, arthropods also have a wide range of chemical and mechanical sensors, mostly based on modifications of the many setae that project through their cuticles. Aquatic species use internal or external fertilization. Almost all arthropods lay eggs, but scorpions give birth to live young after the eggs have hatched inside the mother, arthropod hatchlings vary from miniature adults to grubs and caterpillars that lack jointed limbs and eventually undergo a total metamorphosis to produce the adult form. The level of care for hatchlings varies from nonexistent to the prolonged care provided by scorpions. The evolutionary ancestry of arthropods dates back to the Cambrian period, the group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians in a superphylum Ecdysozoa. Overall however, the relationships of Metazoa are not yet well resolved. Likewise, the relationships between various groups are still actively debated. Arthropods contribute to the food supply both directly as food, and more importantly indirectly as pollinators of crops. Some species are known to spread disease to humans, livestock. The word arthropod comes from the Greek ἄρθρον árthron, joint, and πούς pous, i. e. foot or leg, arthropods are invertebrates with segmented bodies and jointed limbs
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Mollusca
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The molluscs compose the large phylum Mollusca of invertebrate animals. Around 85,000 extant species of molluscs are recognized, molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats and they are highly diverse, not just in size and in anatomical structure, but also in behaviour and in habitat. The phylum is divided into 9 or 10 taxonomic classes. The gastropods are by far the most numerous molluscs in terms of classified species, the three most universal features defining modern molluscs are a mantle with a significant cavity used for breathing and excretion, the presence of a radula, and the structure of the nervous system. Other than these things, molluscs express great morphological diversity, so many textbooks base their descriptions on an ancestral mollusc. This has a single, limpet-like shell on top, which is made of proteins and chitin reinforced with calcium carbonate, the underside of the animal consists of a single muscular foot. Although molluscs are coelomates, the coelom tends to be small, the main body cavity is a hemocoel through which blood circulates, their circulatory systems are mainly open. The generalized mollusc has two paired nerve cords, or three in bivalves, the brain, in species that have one, encircles the esophagus. Most molluscs have eyes, and all have sensors to detect chemicals, vibrations, the simplest type of molluscan reproductive system relies on external fertilization, but more complex variations occur. All produce eggs, from which may emerge trochophore larvae, more complex veliger larvae, good evidence exists for the appearance of gastropods, cephalopods and bivalves in the Cambrian period 541 to 485.4 million years ago. Molluscs have, for centuries, also been the source of important luxury goods, notably pearls, mother of pearl, Tyrian purple dye and their shells have also been used as money in some preindustrial societies. Mollusc species can also represent hazards or pests for human activities, the bite of the blue-ringed octopus is often fatal, and that of Octopus apollyon causes inflammation that can last for over a month. Stings from a few species of large tropical cone shells can also kill, schistosomiasis is transmitted to humans via water snail hosts, and affects about 200 million people. Snails and slugs can also be serious pests, and accidental or deliberate introduction of some snail species into new environments has seriously damaged some ecosystems. The words mollusc and mollusk are both derived from the French mollusque, which originated from the Latin molluscus, from mollis, molluscus was itself an adaptation of Aristotles τα μαλακά, the soft things, which he applied to cuttlefish. The scientific study of molluscs is accordingly called malacology, as it is now known these groups have no relation to molluscs, and very little to one another, the name Molluscoida has been abandoned. The most universal features of the structure of molluscs are a mantle with a significant cavity used for breathing and excretion
28.
Evolutionary history of plants
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The ascendance of flowering plants over gymnosperms in terrestrial environments. Evidence for the appearance of the first land plants occurs in the Ordovician, around 450 million years ago and this chert, formed in volcanic hot springs, preserved several species of early plants in cellular detail by petrification. By the middle of the Devonian, many of the features recognised in plants today were present, including roots, Late Devonian free-sporing plants such as Archaeopteris had secondary vascular tissue that produced wood and had formed forests of tall trees. Also by late Devonian, Elkinsia, an early seed fern, had evolved seeds, evolutionary innovation continued into the Carboniferous and is still ongoing today. Most plant groups were relatively unscathed by the Permo-Triassic extinction event and this may have set the scene for the appearance of the flowering plants in the Triassic, and their later diversification in the Cretaceous and Paleogene. The latest major group of plants to evolve were the grasses, the grasses, as well as many other groups, evolved new mechanisms of metabolism to survive the low CO2 and warm, dry conditions of the tropics over the last 10 million years. Land plants evolved from a group of algae, perhaps as early as 510 million years ago, some molecular estimates place their origin even earlier. Co-operative interactions with fungi may have helped early plants adapt to the stresses of the terrestrial realm and these organisms, although phylogenetically diverse, were probably small and simple, forming little more than an algal scum. The first evidence of plants on land comes from spores of mid-Ordovician age and their walls contain sporopollenin – further evidence of an embryophytic affinity. It could be that atmospheric poisoning prevented eukaryotes from colonising the land prior to this, trilete spores similar to those of vascular plants appear soon afterwards, in Upper Ordovician rocks. Depending exactly when the tetrad splits, each of the four spores may bear a trilete mark, however, this requires that the spore walls be sturdy and resistant at an early stage. This resistance is associated with having a desiccation-resistant outer wall—a trait only of use when spores must survive out of water. Indeed, even those embryophytes that have returned to the lack a resistant wall. The earliest megafossils of land plants were thalloid organisms, which dwelt in fluvial wetlands and are found to have covered most of an early Silurian flood plain and they could only survive when the land was waterlogged. Once plants had reached the land, there were two approaches to dealing with desiccation, the bryophytes avoid it or give in to it, restricting their ranges to moist settings, or drying out and putting their metabolism on hold until more water arrives. Tracheophytes also developed vascular tissue to aid in the movement of water within the organisms, vascular tissue also facilitated upright growth without the support of water and paved the way for the evolution of larger plants on land. The establishment of a land-based flora caused increased accumulation of oxygen in the atmosphere, when this concentration rose above 13%, wildfires became possible. This is first recorded in the early Silurian fossil record by charcoalified plant fossils, apart from a controversial gap in the Late Devonian, charcoal is present ever since
29.
Mammal
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Mammals are any vertebrates within the class Mammalia, a clade of endothermic amniotes distinguished from reptiles by the possession of a neocortex, hair, three middle ear bones and mammary glands. All female mammals nurse their young with milk, secreted from the mammary glands, Mammals include the largest animals on the planet, the great whales. The basic body type is a quadruped, but some mammals are adapted for life at sea, in the air, in trees. The largest group of mammals, the placentals, have a placenta, Mammals range in size from the 30–40 mm bumblebee bat to the 30-meter blue whale. With the exception of the five species of monotreme, all modern mammals give birth to live young, most mammals, including the six most species-rich orders, belong to the placental group. The largest orders are the rodents, bats and Soricomorpha, the next three biggest orders, depending on the biological classification scheme used, are the Primates, the Cetartiodactyla, and the Carnivora. Living mammals are divided into the Yinotheria and Theriiformes There are around 5450 species of mammal, in some classifications, extant mammals are divided into two subclasses, the Prototheria, that is, the order Monotremata, and the Theria, or the infraclasses Metatheria and Eutheria. The marsupials constitute the group of the Metatheria, and include all living metatherians as well as many extinct ones. Much of the changes reflect the advances of cladistic analysis and molecular genetics, findings from molecular genetics, for example, have prompted adopting new groups, such as the Afrotheria, and abandoning traditional groups, such as the Insectivora. The mammals represent the only living Synapsida, which together with the Sauropsida form the Amniota clade, the early synapsid mammalian ancestors were sphenacodont pelycosaurs, a group that produced the non-mammalian Dimetrodon. At the end of the Carboniferous period, this group diverged from the line that led to todays reptiles. Some mammals are intelligent, with some possessing large brains, self-awareness, Mammals can communicate and vocalize in several different ways, including the production of ultrasound, scent-marking, alarm signals, singing, and echolocation. Mammals can organize themselves into fission-fusion societies, harems, and hierarchies, most mammals are polygynous, but some can be monogamous or polyandrous. They provided, and continue to provide, power for transport and agriculture, as well as commodities such as meat, dairy products, wool. Mammals are hunted or raced for sport, and are used as model organisms in science, Mammals have been depicted in art since Palaeolithic times, and appear in literature, film, mythology, and religion. Defaunation of mammals is primarily driven by anthropogenic factors, such as poaching and habitat destruction, Mammal classification has been through several iterations since Carl Linnaeus initially defined the class. No classification system is accepted, McKenna & Bell and Wilson & Reader provide useful recent compendiums. Though field work gradually made Simpsons classification outdated, it remains the closest thing to a classification of mammals
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Flowering plant
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The flowering plants, also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants, with 416 families, approx. 13,164 known genera and a total of c.295,383 known species, etymologically, angiosperm means a plant that produces seeds within an enclosure, in other words, a fruiting plant. The term angiosperm comes from the Greek composite word meaning enclosed seeds, the ancestors of flowering plants diverged from gymnosperms in the Triassic Period, during the range 245 to 202 million years ago, and the first flowering plants are known from 160 mya. They diversified extensively during the Lower Cretaceous, became widespread by 120 mya, angiosperms differ from other seed plants in several ways, described in the table. These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants, the amount and complexity of tissue-formation in flowering plants exceeds that of gymnosperms. The vascular bundles of the stem are arranged such that the xylem and phloem form concentric rings, in the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium, the soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Among the monocotyledons, the bundles are more numerous in the stem and are scattered through the ground tissue. They contain no cambium and once formed the stem increases in diameter only in exceptional cases, the characteristic feature of angiosperms is the flower. Flowers show remarkable variation in form and elaboration, and provide the most trustworthy external characteristics for establishing relationships among angiosperm species, the function of the flower is to ensure fertilization of the ovule and development of fruit containing seeds. The floral apparatus may arise terminally on a shoot or from the axil of a leaf, occasionally, as in violets, a flower arises singly in the axil of an ordinary foliage-leaf. There are two kinds of cells produced by flowers. Microspores, which divide to become pollen grains, are the male cells and are borne in the stamens. The female cells called megaspores, which divide to become the egg cell, are contained in the ovule. The flower may consist only of parts, as in willow. Usually, other structures are present and serve to protect the sporophylls, the individual members of these surrounding structures are known as sepals and petals. The outer series is usually green and leaf-like, and functions to protect the rest of the flower, the inner series is, in general, white or brightly colored, and is more delicate in structure. It functions to attract insect or bird pollinators, attraction is effected by color, scent, and nectar, which may be secreted in some part of the flower
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Primate
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A primate is a mammal of the order Primates. In taxonomy, primates include two distinct lineages, strepsirrhines and haplorhines, Primates arose from ancestors that lived in the trees of tropical forests, many primate characteristics represent adaptations to life in this challenging three-dimensional environment. Most primate species remain at least partly arboreal, with the exception of humans, who inhabit every continent except for Antarctica, most primates live in tropical or subtropical regions of the Americas, Africa and Asia. Based on fossil evidence, the earliest known true primates, represented by the genus Teilhardina, an early close primate relative known from abundant remains is the Late Paleocene Plesiadapis, c. Molecular clock studies suggest that the branch may be even older. The order Primates was traditionally divided into two groupings, prosimians and anthropoids. Prosimians have characteristics more like those of the earliest primates, and include the lemurs of Madagascar, lorisoids, simians include monkeys, apes and hominins. Simians are divided into two groups, catarrhine monkeys and apes of Africa and Southeast Asia and platyrrhine or New World monkeys of South, catarrhines consist of Old World monkeys, gibbons and great apes, New World monkeys include the capuchin, howler and squirrel monkeys. Humans are the only extant catarrhines to have spread successfully outside of Africa, South Asia, New primate species are still being discovered. More than 25 species were described in the decade of the 2000s. Considered generalist mammals, primates exhibit a range of characteristics. Some primates are primarily terrestrial rather than arboreal, but all species possess adaptations for climbing trees, locomotion techniques used include leaping from tree to tree, walking on two or four limbs, knuckle-walking, and swinging between branches of trees. Primates are characterized by large brains relative to other mammals, as well as a reliance on stereoscopic vision at the expense of smell. These features are developed in monkeys and apes and noticeably less so in lorises. Three-color vision has developed in some primates, most also have opposable thumbs and some have prehensile tails. Many species are dimorphic, differences include body mass, canine tooth size. Primates have slower rates of development than other similarly sized mammals and reach maturity later, depending on the species, adults may live in solitude, in mated pairs, or in groups of up to hundreds of members. The relationships among the different groups of primates were not clearly understood until relatively recently, for example, ape has been used either as an alternative for monkey or for any tailless, relatively human-like primate
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History of Earth
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The history of Earth concerns the development of the planet Earth from its formation to the present day. Nearly all branches of science have contributed to the understanding of the main events of the Earths past. The age of Earth is approximately one-third of the age of the universe, an immense amount of geological change has occurred in that timespan, accompanied by the emergence of life and its subsequent evolution. Earth formed around 4.54 billion years ago by accretion from the solar nebula, volcanic outgassing probably created the primordial atmosphere and then the ocean, but the atmosphere contained almost no oxygen and so would have been toxic to most modern life including humans. Much of the Earth was molten because of frequent collisions with other bodies which led to extreme volcanism. A giant impact collision with a body named Theia while Earth was in its earliest stage. Over time, the Earth cooled, causing the formation of a solid crust, the geological time scale clock depicts the larger spans of time from the beginning of the Earth as well as a chronology of some definitive events of Earth history. The Archean and Proterozoic eons follow, they produced the abiogenesis of life on Earth, there are microbial mat fossils such as stromatolites found in 3.48 billion-year-old sandstone discovered in Western Australia. According to one of the researchers, If life arose relatively quickly on Earth … then it could be common in the universe, living forms derived from photosynthesis appeared between 3.2 and 2.4 billion years ago and began enriching the atmosphere with oxygen. More than 99 percent of all species, amounting to five billion species. Estimates on the number of Earths current species range from 10 million to 14 million, of which about 1.2 million have been documented and over 86 percent have not yet been described. More recently, in May 2016, scientists reported that 1 trillion species are estimated to be on Earth currently with only one-thousandth of one percent described. Geological change has been a constant of Earths crust since the time of its formation, species continue to evolve, taking on new forms, splitting into daughter species or going extinct in the process of adapting or dying in response to ever-changing physical environments. The process of plate tectonics continues to play a dominant role in the shaping of Earths oceans and continents, in geochronology, time is generally measured in mya, each unit representing the period of approximately 1,000,000 years in the past. The history of Earth is divided into four great eons, starting 4,540 mya with the formation of the planet, each eon saw the most significant changes in Earths composition, climate and life. Each eon is divided into eras, which in turn are divided into periods. The history of the Earth can be organized according to the geologic time scale. The following four timelines show the time scale
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Age of the Earth
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The age of the Earth is 4.54 ±0.05 billion years. This dating is based on evidence from radiometric age-dating of meteorite material and is consistent with the ages of the oldest-known terrestrial. Following the development of radiometric age-dating in the early 20th century, the oldest such minerals analyzed to date—small crystals of zircon from the Jack Hills of Western Australia—are at least 4.404 billion years old. Comparing the mass and luminosity of the Sun to those of other stars and it is hypothesised that the accretion of Earth began soon after the formation of the calcium-aluminium-rich inclusions and the meteorites. It is also difficult to determine the age of the oldest rocks on Earth, exposed at the surface. Studies of strata, the layering of rocks and earth, gave naturalists an appreciation that Earth may have been many changes during its existence. These layers often contained fossilized remains of creatures, leading some to interpret a progression of organisms from layer to layer. Nicolas Steno in the 17th century was one of the first naturalists to appreciate the connection between fossil remains and strata and his observations led him to formulate important stratigraphic concepts. In the 1790s, William Smith hypothesized that if two layers of rock at widely differing locations contained similar fossils, then it was plausible that the layers were the same age. William Smiths nephew and student, John Phillips, later calculated by means that Earth was about 96 million years old. In the mid-18th century, the naturalist Mikhail Lomonosov suggested that Earth had been created separately from, and several hundred years before. In 1779 the Comte du Buffon tried to obtain a value for the age of Earth using an experiment, He created a globe that resembled Earth in composition. This led him to estimate that Earth was about 75,000 years old, other naturalists used these hypotheses to construct a history of Earth, though their timelines were inexact as they did not know how long it took to lay down stratigraphic layers. This was a challenge to the view, which saw the history of Earth as static. Many naturalists were influenced by Lyell to become uniformitarians who believed that changes were constant, in 1862, the physicist William Thomson, 1st Baron Kelvin published calculations that fixed the age of Earth at between 20 million and 400 million years. He assumed that Earth had formed as a completely molten object, geologists such as Charles Lyell had trouble accepting such a short age for Earth. For biologists, even 100 million years seemed much too short to be plausible, in Darwins theory of evolution, the process of random heritable variation with cumulative selection requires great durations of time. In a lecture in 1869, Darwins great advocate, Thomas H. Huxley, attacked Thomsons calculations and their values were consistent with Thomsons calculations
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Origin of water on Earth
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The origin of water on Earth, or the reason that there is clearly more liquid water on Earth than on the other rocky planets of the Solar System, is not completely understood. There exist numerous more or less mutually compatible hypotheses as to how water may have accumulated on Earths surface over the past 4.5 billion years in sufficient quantity to form oceans. Comets, trans-Neptunian objects or water-rich meteoroids from the reaches of the asteroid belt colliding with Earth may have brought water to the worlds oceans. Large enough Planetesimals were heated by the decay of aluminium isotope and this could cause water to rise to the surface. What is however unclear is whether these comets are representative of those from the Kuiper Belt, the water in carbon-rich chondrites point to a similar D/H ratio as oceanic water. Nevertheless, mechanisms have been proposed to suggest that the D/H-ratio of oceanic water may have increased significantly throughout Earths history, such a proposal is consistent with the possibility that a significant amount of the water on Earth was already present during the planets early evolution. Recent measurements of the composition of Moon rocks suggest that Earth was born with its water already present. Investigating lunar samples carried to Earth by the Apollo 15 and 17 missions found a ratio that matched the isotopic ratio in carbonaceous chondrites. The ratio is similar to that found in water on Earth. The findings suggest a common source of water for both objects and this supports a theory that Jupiter temporarily migrated into the inner Solar System, destabilizing the orbits of water-rich carbonaceous chondrites. As a result, some of the bodies could have fallen inwards and become part of the raw material for making Earth, the discovery of water vapor out-gassing from Ceres provides related information on water-ice content of the asteroid belt. Gradual leakage of stored in hydrate minerals of Earths rocks could have formed a portion of its water. Water may also have come from volcanism, water vapor in the atmosphere that originated in volcanic eruptions may have condensed to form rain, a sizeable quantity of water would have been in the material that formed Earth. Water molecules would have escaped Earths gravity more easily when it was less massive during its formation, part of the young planet is theorized to have been disrupted by the impact which created the Moon, which should have caused melting of one or two large areas. Present composition does not match complete melting and it is hard to melt, however, a fair fraction of material should have been vaporized by this impact, creating a rock-vapor atmosphere around the young planet. The rock-vapor would have condensed within two years, leaving behind hot volatiles which probably resulted in a heavy carbon dioxide atmosphere with hydrogen. Liquid water oceans existed despite the temperature of 230°C because of the atmospheric pressure of the heavy CO2 atmosphere. As cooling continued, subduction and dissolving in ocean water removed most CO2 from the atmosphere but levels oscillated wildly as new surface and mantle cycles appeared
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Earliest known life forms
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The earliest known life forms on Earth are putative fossilized microorganisms. The earliest time that life forms first appeared on Earth is unknown, a life form, or lifeform, is an entity or being that is living. Earth remains the place in the universe known to harbor life forms. More than 99% of all species of life forms, amounting to five billion species. Estimates on the number of Earths current species of life forms range from 10 million to 14 million, of which about 1.2 million have been documented and over 86 percent have not yet been described. More recently, in May 2016, scientists reported that 1 trillion species are estimated to be on Earth currently with only one-thousandth of one percent described, the total amount of related DNA base pairs on Earth is estimated at 5.0 x 1037 and weighs 50 billion tonnes. In comparison, the mass of the biosphere has been estimated to be as much as 4 TtC. In July 2016, scientists reported identifying a set of 355 genes from the Last Universal Common Ancestor of all living on Earth. Under certain test conditions, life forms have been observed to thrive in the vacuum of outer space, Life forms appear to thrive in the Mariana Trench, the deepest spot in the Earths oceans. In August 2014, scientists confirmed the existence of life forms living 800 m below the ice of Antarctica, fossil evidence from the Earth informs most studies of the origin of life. Microbial mat fossils have found in 3.48 billion-year-old sandstone in Western Australia. Other early physical evidence of biogenic substances includes graphite, and possibly stromatolites, remains of biotic life have been found in 4.1 billion-year-old rocks in Western Australia. In recent years, there have been a number of discoveries that suggested the earliest appearance of life on Earth was even earlier in time, according to biologist Stephen Blair Hedges, If life arose relatively quickly on Earth … then it could be common in the universe. Biota Life Vitae Wikispecies – a free directory of life
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Late Heavy Bombardment
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The Late Heavy Bombardment is an event thought to have occurred approximately 4.1 to 3.8 billion years ago, at a time corresponding to the Neohadean and Eoarchean eras on Earth. The LHB happened after the Earth and other rocky planets had formed and accreted most of their mass, Evidence for the LHB derives from lunar samples brought back by the Apollo astronauts. Isotopic dating of Moon rocks implies that most impact melts occurred in a narrow interval of time. Several hypotheses attempt to explain the apparent spike in the flux of impactors in the inner Solar System and they also note that the rate of impact cratering could differ significantly between the outer and inner zones of the Solar System. The main piece of evidence for a lunar cataclysm comes from the ages of impact melt rocks that were collected during the Apollo missions. The majority of impact melts are believed to have formed during the collision of asteroids or comets tens of kilometers across. The Apollo 15,16, and 17 landing sites were chosen as a result of their proximity to the Imbrium, Nectaris, under study on Earth, the ages of impact melts collected at these sites clustered between about 3.8 and 4.1 Ga. The apparent clustering of ages of these led to postulation that the record an intense bombardment of the Moon. They called it the lunar cataclysm and proposed that it represented an increase in the rate of bombardment of the Moon around 3.9 Ga. At the time, the conclusion was considered controversial, as more data has become available, particularly from lunar meteorites, this theory, while still controversial, has gained in popularity. The lunar meteorites are believed to randomly sample the lunar surface, many of the feldspathic lunar meteorites probably originated from the lunar far side, and impact melts within these have recently been dated. Consistent with the hypothesis, none of their ages was found to be older than about 3.9 Ga. Nevertheless, the ages do not cluster at this date. Dating of howardite, eucrite and diogenite meteorites and H chondrite meteorites originating from the asteroid belt reveal numerous ages from 3. 4–4.1 Ga, impact velocities above 10 km/s require very high inclinations or the large eccentricities of asteroids on planet crossing orbits. Such objects are rare in the current asteroid belt but the population would be increased by the sweeping of resonances due to giant planet migration. Studies of the highland crater size distributions suggest that the family of projectiles struck Mercury. The first criticism concerns the origin of the impact melt rocks that were sampled at the Apollo landing sites. While these impact melts have been attributed to having been derived from the closest basin. According to this hypothesis, the cluster of impact melt ages near 3.9 Ga simply reflects material being collected from a single impact event, Imbrium
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Geological history of oxygen
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Before photosynthesis evolved, Earths atmosphere had no free oxygen. Photosynthetic prokaryotic organisms that produced O2 as a waste product lived long before the first build-up of free oxygen in the atmosphere, the oxygen they produced would have been rapidly removed from the atmosphere by weathering of reducing minerals, most notably iron. This mass rusting led to the deposition of oxide on the ocean floor. Oxygen only began to persist in the atmosphere in small quantities about 50 million years before the start of the Great Oxygenation Event and this mass oxygenation of the atmosphere resulted in rapid buildup of free oxygen. At current rates of production, todays concentration of oxygen could be produced by photosynthetic organisms in 2,000 years.9 billion years ago. The presence of O2 provided life with new opportunities, aerobic metabolism is more efficient than anaerobic pathways, and the presence of oxygen undoubtedly created new possibilities for life to explore. Since the start of the Cambrian period, atmospheric oxygen concentrations have fluctuated between 15% and 35% of atmospheric volume. The maximum of 35% was reached towards the end of the Carboniferous period, whilst human activities, such as the burning of fossil fuels, affect relative carbon dioxide concentrations, their effect on the much larger concentration of oxygen is less significant. But Haldanes essay points out that it would apply to insects. However, the basis for this correlation is not firm. There is no significant correlation between atmospheric oxygen and maximum body size elsewhere in the geological record, ecological constraints can better explain the diminutive size of post-Carboniferous dragonflies - for instance, the appearance of flying competitors such as pterosaurs, birds and bats. The most celebrated link between oxygen and evolution occurs at the end of the last of the Snowball glaciations, where complex life is first found in the fossil record. Significant concentrations of oxygen were just one of the prerequisites for the evolution of complex life, models based on uniformitarian principles suggest that such a concentration was only reached immediately before metazoa first appeared in the fossil record. First breath, Earths billion-year struggle for oxygen New Scientist, #2746,5 February 2010 by Nick Lane, the Mystery of Earths Oxygen New York Times,3 October 2013 by Carl Zimmer
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Great Oxygenation Event
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The Great Oxygenation Event was the biologically induced appearance of dioxygen in Earths atmosphere. The current geochemical and biomarker evidence for the development of oxygenic photosynthesis before the Great Oxidation Event has been mostly inconclusive. Oceanic cyanobacteria, which evolved into multicellular forms more than 2.3 billion years ago, are believed to have become the first microbes to produce oxygen by photosynthesis, before the GOE, any free oxygen they produced was chemically captured by dissolved iron or organic matter. The GOE was the point in time when these oxygen sinks became saturated, at which point oxygen, the increased production of oxygen set Earths original atmosphere off balance. Free oxygen is toxic to anaerobic organisms, and the rising concentrations may have destroyed most such organisms at the time. Cyanobacteria were therefore responsible for one of the most significant extinction events in Earths history, besides marine cyanobacteria, there is also evidence of cyanobacteria on land. A spike in chromium contained in ancient rock deposits formed underwater shows they had accumulated chromium washed off from continental shelves, chromium is not easily dissolved and its release from rocks would have required the presence of a powerful acid. One such acid, sulfuric acid, might have created through bacterial reactions with pyrite. Eventually, the evolution of organisms that consumed oxygen established an equilibrium in its availability. Free oxygen has been an important constituent of the atmosphere ever since, the excess of oxygen those organisms produce is a waste product. These organisms lived long before the GOE, perhaps as early as 3,500 million years ago, initially, the oxygen they produced would have quickly been removed from the atmosphere by the chemical weathering of reducing minerals, most notably iron. This mass rusting led to the deposition of oxide in the form of banded-iron formations such as the sediments in Minnesota and Pilbara. Oxygen only began to persist in the atmosphere in small quantities shortly before the start of the GOE, without a mineral sink in the form of iron, oxygen could have accumulated very rapidly. For example, at rates of photosynthesis, modern atmospheric O2 levels could be produced in around 2,000 years. Another hypothesis is that producers did not evolve until just before the major rise in atmospheric oxygen concentration. This is based on interpretation of the oxygen indicator, mass-independent fractionation of sulfur isotopes. This hypothesis would eliminate the need to explain a lag in time between the evolution of oxyphotosynthetic microbes and the rise in free oxygen.4 to 2.1 billion years ago. The latter may have been a full-blown, and possibly the longest ever, snowball Earth episode, despite the natural recycling of organic matter, life had remained energetically limited until the widespread availability of oxygen
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Ediacaran biota
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The Ediacaran biota consisted of enigmatic tubular and frond-shaped, mostly sessile organisms that lived during the Ediacaran Period. Trace fossils of organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The Ediacaran biota radiated in an event called the Avalon explosion,575 million years ago, the biota largely disappeared with the rapid increase in biodiversity known as the Cambrian explosion. Most of the existing body plans of animals first appeared in the fossil record of the Cambrian rather than the Ediacaran. For macroorganisms, the Cambrian biota appears to have replaced the organisms that dominated the Ediacaran fossil record. Multiple hypotheses exist to explain the disappearance of this biota, including bias, a changing environment. The morphology and habit of some taxa suggest relationships to Porifera or Cnidaria, Kimberella may show a similarity to molluscs, and other organisms have been thought to possess bilateral symmetry, although this is controversial. Most macroscopic fossils are morphologically distinct from later life-forms, they resemble discs, tubes, one palaeontologist proposed a separate kingdom level category Vendozoa in the Linnaean hierarchy for the Ediacaran biota. The concept of Ediacaran Biota is, of course, somewhat artificial as it can not be defined geographically, stratigraphically, taphonomically, the first Ediacaran fossils discovered were the disc-shaped Aspidella terranovica in 1868. Their discoverer, Scottish geologist Alexander Murray, found them useful aids for correlating the age of rocks around Newfoundland, instead, they were interpreted as gas escape structures or inorganic concretions. No similar structures elsewhere in the world were known and the one-sided debate soon fell into obscurity. It was not until the British discovery of the iconic Charnia in 1957 that the pre-Cambrian was seriously considered as containing life. This frond-shaped fossil was found in Englands Charnwood Forest, and due to the geological mapping of the British Geological Survey there was no doubt these fossils sat in Precambrian rocks. All specimens discovered until 1967 were in coarse-grained sandstone that prevented preservation of fine details, poor communication, combined with the difficulty in correlating globally distinct formations, led to a plethora of different names for the biota. Ediacaran and Ediacarian were subsequently applied to the epoch or period of geological time, in March 2004, the International Union of Geological Sciences ended the inconsistency by formally naming the terminal period of the Neoproterozoic after the Australian locality. The term Ediacaran biota and similar has, at times, been used in a geographic, stratigraphic, taphonomic, or biological sense. Microbial mats are areas of sediment stabilised by the presence of colonies of microbes that secrete sticky fluids or otherwise bind the sediment particles. They appear to migrate upwards when covered by a layer of sediment but this is an illusion caused by the colonys growth, individuals do not, themselves
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Cambrian explosion
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Lasting for about the next 20–25 million years, it resulted in the divergence of most modern metazoan phyla. Additionally, the event was accompanied by major diversification of other organisms, prior to the Cambrian explosion, most organisms were simple, composed of individual cells occasionally organized into colonies. Over the following 70 to 80 million years, the rate of diversification accelerated by an order of magnitude, almost all present animal phyla appeared during this period. There is strong evidence for species of Cnidaria and Porifera existing in the Ediacaran, bryozoans dont appear in the fossil record until after the Cambrium, in the Lower Ordovician. The Cambrian explosion has generated extensive scientific debate, interpretation is difficult due to a limited supply of evidence, based mainly on an incomplete fossil record and chemical signatures remaining in Cambrian rocks. The first discovered Cambrian fossils were trilobites, described by Edward Lhuyd, nineteenth-century geologists such as Adam Sedgwick and Roderick Murchison used the fossils for dating rock strata, specifically for establishing the Cambrian and Silurian periods. He reasoned that earlier seas had swarmed with living creatures, earlier fossil evidence has since been found. Fossils of more complex cells, from which all animals, plants. Rocks dating from 580 to 543 million years ago contain fossils of the Ediacara biota, organisms so large that they are likely multicelled, the most common organism, Marrella, was clearly an arthropod, but not a member of any known arthropod class. Stephen Jay Goulds popular 1989 account of work, Wonderful Life, brought the matter into the public eye. While differing significantly in details, both Whittington and Gould proposed that all animal phyla had appeared almost simultaneously in a rather short span of geological period. Other analyses, some recent and some dating back to the 1970s. Radiometric dates for much of the Cambrian, obtained by analysis of elements contained within rocks, have only recently become available. Therefore, dates or descriptions of sequences of events should be regarded with some caution until better data become available, fossils of organisms bodies are usually the most informative type of evidence. Fossilization is an event, and most fossils are destroyed by erosion or metamorphism before they can be observed. Hence, the record is very incomplete, increasingly so as earlier times are considered. Despite this, they are adequate to illustrate the broader patterns of lifes history. Also, biases exist in the record, different environments are more favourable to the preservation of different types of organism or parts of organisms
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Tetrapod
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The superclass Tetrapoda contains the four-limbed vertebrates known as tetrapods, it includes living and extinct amphibians, reptiles and mammals, as well as earlier extinct groups. The first tetrapods appeared by the late Devonian,367, the first tetrapods were primarily aquatic. Their descendants drove most amphibians to extinction as they began to dominate, the amniotes include the tetrapods that further evolved for flight—such as birds from among the dinosaurs, and bats from among the mammals. The change from a plan for breathing and navigating in water to a body plan enabling the animal to move on land is one of the most profound evolutionary changes known. It is also becoming increasingly well-understood as a result of more transitional fossil finds, many tetrapods have returned to partially aquatic or fully aquatic lives throughout the history of the group. Tetrapods can be defined in cladistics as the nearest common ancestor of all living amphibians and all living amniotes, the group so defined is the crown group, or crown tetrapods. The term tetrapodomorph is used for the definition, any animal that is more closely related to living amphibians, reptiles, birds. The group so defined is known as the total group. Stegocephalia is a larger group equivalent to some uses of the word tetrapod. Batrachomorphs are all sharing a more recent common ancestry with living amphibians than with living amniotes. Reptiliomorphs are all sharing a more recent common ancestry with living amniotes than with living amphibians. Tetrapoda includes four classes, amphibians, reptiles, mammals, however, that diversification process was interrupted at least a few times by major biological crises, such as the Permian–Triassic extinction event, which at least affected amniotes. As biodiversity has grown, so has the number of niches that tetrapods have occupied, the first tetrapods were aquatic and fed primarily on fish. Today, the Earth supports a diversity of tetrapods that live in many habitats. Tetrapods evolved from early bony fishes, specifically from the branch of lobe-finned fishes. The first tetrapods probably evolved in the Emsian stage of the Early Devonian from Tetrapodomorph fish living in water environments. The very earliest tetrapods would have been similar to Acanthostega, with legs and lungs as well as gills. The earliest tetrapods inhabited saltwater, brackish-water, and freshwater environments and these traits were shared with many early lobed-finned fishes
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Ape
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The simians are the monkeys, cladistically including the apes, the New World monkeys or platyrrhines, and the catarrhine clade consisting of the Old World monkeys and apes. The simian line and the line diverged about 60 million years ago. Forty million years ago, simians from Africa colonized South America, the remaining simians split 25 million years ago into apes and Old World monkeys. Under modern classification, the tarsiers and simians are grouped under the suborder Haplorhini while the strepsirrhines are placed in suborder Strepsirrhini, in anthropoidea, evidences indicate that the Old and the New World primates went through parallel evolution. Primatology, paleoanthropology, and other related fields are split on their usage of the synonymous infraorder names, Simiiformes and Anthropoidea. According to Robert Hoffstetter, the term Simiiformes has priority over Anthropoidea because of the taxonomic term Simii by van der Hoeven, from which it is constructed, dates to 1833. In contrast, Anthropoidea by Mivart dates to 1864, while Simiiformes by Haeckel dates to 1866, Hoffstetter also argued that Simiiformes is also constructed like a proper infraorder name, whereas Anthropoidea ends in -oidea, which is reserved for superfamilies. He also noted that Anthropoidea is too easily confused with anthropoïdes, the simians are split into three distinct groups. The New World monkeys in parvorder Platyrrhini split from the rest of the line about 40 mya. This group split about 25 mya between the Old World monkeys and the apes
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Phanerozoic
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The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 541 million years to the present, and began with the Cambrian Period when diverse hard-shelled animals first appeared. Its name was derived from the Ancient Greek words φανερός and ζωή, meaning life, since it was once believed that life began in the Cambrian. The time before the Phanerozoic, called the Precambrian supereon, is now divided into the Hadean, Archaean, land plant life appeared in the early Phanerozoic eon. During this time span, tectonic forces caused the continents to move and eventually collect into a landmass known as Pangaea. The Proterozoic-Phanerozoic boundary is at 541 million years ago, the three different dividing points are within a few million years of each other. The Phanerozoic is divided into three eras, the Paleozoic, Mesozoic, and Cenozoic, which are subdivided into 12 periods. The Paleozoic features the rise of fish, amphibians and reptiles, the Mesozoic is ruled by the reptiles, and features the evolution of mammals, birds and more famously, dinosaurs. The Cenozoic is the time of the mammals, and more recently, the Paleozoic is a time in Earths history when complex life forms evolved, took their first breath of oxygen on dry land, and when the forerunners of all life on Earth began to diversify. There are six periods in the Paleozoic era, Cambrian, Ordovician, Silurian, Devonian, Carboniferous, the Cambrian is the first period of the Paleozoic Era and starts from 541 to 485 million years ago. The Cambrian sparked a rapid expansion in evolution in an event known as the Cambrian Explosion during which the greatest number of creatures evolved in a period in the history of Earth. Plants like algae evolved, and the fauna was dominated by armored arthropods, almost all marine phyla evolved in this period. During this time, the super-continent Pannotia began to break up, the Ordovician spans from 485 million years to 440 million years. The Ordovician was a time in Earths history in many species still prevalent today evolved, such as primitive fish, cephalopods. The most common forms of life, however, were trilobites, snails, more importantly, the first arthropods crept ashore to colonize Gondwana, a continent empty of animal life. By the end of the Ordovican, Gondwana had moved from the equator to the South Pole, the glaciation of Gondwana resulted in a major drop in sea level, killing off all life that had established along its coast. Glaciation caused a snowball Earth, leading to the Ordovician-Silurian extinction, during which 60% of marine invertebrates and this is considered the first mass extinction and the second deadliest in the history of Earth. The Silurian spans from 440 million years to 415 million years, fully terrestrial life evolved, which included early arachnids, fungi, and centipedes
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Proterozoic
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The Proterozoic is a geological eon representing the time just before the proliferation of complex life on Earth. The name Proterozoic comes from Greek and means life, the Greek root protero-, means former, earlier and zoic-, means animal. The Proterozoic Eon extended from 2500 Ma to 541 Ma, and is the most recent part of the Precambrian Supereon and it is subdivided into three geologic eras, the Paleoproterozoic, Mesoproterozoic, and Neoproterozoic. The geologic record of the Proterozoic Eon is more complete than that for the preceding Archean Eon, studies of these rocks have shown that the eon continued the massive continental accretion that had begun late in the Archean Eon. The Proterozoic Eon also featured the first definitive supercontinent cycles and wholly modern mountain building activity, there is evidence that the first known glaciations occurred during the Proterozoic. One of the most important events of the Proterozoic was the accumulation of oxygen in the Earths atmosphere, until roughly 2.3 billion years ago, oxygen was probably only 1% to 2% of its current level. The Banded iron formations, which provide most of the iron ore, are one mark of that mineral sink process. Their accumulation ceased after 1.9 billion years ago, after the iron in the oceans had all been oxidized, red beds, which are colored by hematite, indicate an increase in atmospheric oxygen 2 billion years ago. Such massive iron oxide formations are not found in older rocks, the Proterozoic Eon was a very tectonically active period in the Earth’s history. The late Archean Eon to Early Proterozoic Eon corresponds to a period of increasing crustal recycling, suggesting subduction, evidence for this increased subduction activity comes from the abundance of old granites originating mostly after 2.6 Ga. The appearance of eclogites, which metamorphic rocks created by pressure, are explained using a model that incorporates subduction. As a result of remelting of basaltic oceanic crust due to subduction, the long-term tectonic stability of those cratons is why we find continental crust ranging up to a few billion years in age. It is believed that 43% of modern continental crust was formed in the Proterozoic, 39% formed in the Archean, studies by Condie 2000 and Rino et al.2004 suggest that crust production happened episodically. By isotopically calculating the ages of Proterozoic granitoids it was determined there were several episodes of rapid increase in continental crust production. The reason for these pulses is unknown, but they seemed to have decreased in magnitude after every period, evidence of collision and rifting between continents raises the question as to what exactly were the movements of the Archean cratons composing Proterozoic continents. Paleomagnetic and geochronological dating mechanisms have allowed the deciphering of Precambrian Supereon tectonics and it is known that tectonic processes of the Proterozoic Eon resemble greatly the evidence of tectonic activity, such as orogenic belts or ophiolite complexes, we see today. Hence, most geologists would conclude that the Earth was active at that time and it is also commonly accepted that during the Precambrian, the Earth went through several supercontinent breakup and rebuilding cycles. In the late Proterozoic, the dominant supercontinent was Rodinia and it consisted of a series of continents attached to a central craton that forms the core of the North American Continent called Laurentia
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Hadean
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The Hadean is a geologic eon of the Earth predating the Archean. It began with the formation of the Earth about 4.6 billion years ago and ended, the geologist Preston Cloud coined the term in 1972, originally to label the period before the earliest-known rocks on Earth. W. Brian Harland later coined an almost synonymous term, the Priscoan period, other, older texts simply refer to the eon as the Pre-Archean. Nonetheless, in 2015, traces of carbon minerals interpreted as remains of life were found in 4. 1-billion-year-old rocks in Western Australia. Since few geological traces of this eon remain on Earth, there is no official subdivision, however, the Lunar geologic timescale embraces several major divisions relating to the Hadean, so these are sometimes used in an informal sense to refer to the same periods of time on Earth. In 2010, a scale was proposed that includes the addition of the Chaotian and Prenephelean Eons preceding the Hadean. The Paleohadean era consists of the Hephaestean and the Jacobian periods, the Mesohadean is divided into the Canadian and the Procrustean periods. The Neohadean is divided into the Acastan and the Promethean periods, as of February 2017, this has not been adopted by the IUGS. In the last decades of the 20th century geologists identified a few Hadean rocks from Western Greenland, Northwestern Canada, and Western Australia. The oldest dated zircon crystals, enclosed in a metamorphosed sandstone conglomerate in the Jack Hills of the Narryer Gneiss Terrane of Western Australia, date to 4.404 ±0.008 Ga. This zircon is an outlier, with the oldest consistently-dated zircon falling closer to 4.35 Ga—around 200 million years after the hypothesized time of the Earths formation. A sizeable quantity of water would have been in the material formed the Earth. Water molecules would have escaped Earths gravity more easily when it was less massive during its formation, hydrogen and helium are expected to continually escape due to atmospheric escape. Part of the ancient planet is theorized to have been disrupted by the impact created the Moon. Earths present composition suggests that there was not complete remelting as it is difficult to completely melt, however, a fair fraction of material should have been vaporized by this impact, creating a rock vapor atmosphere around the young planet. The rock vapor would have condensed within two years, leaving behind hot volatiles which probably resulted in a heavy CO2 atmosphere with hydrogen. Liquid water oceans existed despite the temperature of 230 °C because at an atmospheric pressure of above 27 atmospheres, caused by the heavy CO2 atmosphere. As cooling continued, subduction and dissolving in ocean water removed most CO2 from the atmosphere but levels oscillated wildly as new surface and mantle cycles appeared
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