The black-tailed jackrabbit known as the American desert hare, is a common hare of the western United States and Mexico, where it is found at elevations from sea level up to 10,000 ft. Reaching a length around 2 ft, a weight from 3 to 6 lb, the black-tailed jackrabbit is the third-largest North American hare. Black-tailed jackrabbits occupy mixed shrub-grassland terrains, their breeding depends on the location. Young are born furred with eyes open; the average litter size is around four, but may be as low as two and as high as seven in warm regions. The black-tailed jackrabbit does not migrate or hibernate during winter and uses the same habitat of 0.4 to 1.2 mi2 year-round. Its diet is composed of various shrubs, small trees and forbs. Shrubs comprise the bulk of fall and winter diets, while grasses and forbs are used in spring and early summer, but the pattern and plant species vary with climate; the black-tailed jackrabbit is an important prey species for raptors and carnivorous mammals, such as eagles, owls, coyotes and wild cats.
The rabbits host many ectoparasites including fleas, ticks and mites. Like other jackrabbits, the black-tailed jackrabbit has distinctive long ears, the long powerful rear legs characteristic of hares. Reaching a length about 2 ft, a weight from 3 to 6 lb, the black-tailed jackrabbit is the third-largest North American hare, after the antelope jackrabbit and the white-tailed jackrabbit; the black-tailed jackrabbit's dorsal fur is agouti, its undersides and the insides of its legs are creamy white. The ears are black-tipped on the outer surfaces, unpigmented inside; the ventral surface of the tail is grey to white, the black dorsal surface of the tail continues up the spine for a few inches to form a short, black stripe. The females are larger with no other significant differences. Although 17 subspecies are recognized, this number may be excessive. Using a cluster analysis of anatomical characters and others found that black-tailed jackrabbit subspecies separated into two distinct groups that are geographically separated west and east of the Colorado Rocky Mountains and the Colorado River.
They suggested only two infrataxa are warranted: the western subspecies L. c. californicus and the eastern subspecies L. c. texianus. The black-tailed jackrabbit is the most distributed jackrabbit in North America. Native black-tailed jackrabbit populations occur from central Washington east to Missouri and south to Baja California Sur and Zacatecas. Black-tailed jackrabbit distribution is expanding eastward in the Great Plains at the expense of white-tailed jackrabbit; the black-tailed jackrabbit has been introduced in southern Florida and along the coastline in Maryland, New Jersey, Virginia. In the Baja California Peninsula, there are six subspecies of L. californicus, three of which are endemic to the surrounding islands. The current distribution is a result from sea level rise about 21,000 years ago, after the Last Glacial Maximum. Due to this geographic isolation, the current subspecies of L. californicus living in the peninsula can be separated into 3 subclades based on similar DNA structure and pelage color.
The first clade is associated with subspecies L. c. xanti, contains all subspecies found in the southernmost part of the Baja Peninsula. The second clade is associated with subspecies L. c. magdalenae, includes all subspecies found between the La Paz isthmus and the southern Vixcaino Desert, including subspecies L. c. xanti, L. c. sheldoni, L. c. martirensis. This clade has a ranging coloration pattern of light brown to yellow; the third clade is associated with subspecies L. c. martirensis, includes all subspecies found from the Viscaino Desert to the northernmost part of the peninsula. Distribution of subspecies occurring or in the United States is: Lepus californicus altamirae L. c. asellus L. c. bennettii – coastal southern California to Baja California Norte L. c. californicus – coastal Oregon to coastal and Central Valley California L. c. curti L. c. deserticola – southern Idaho to Sonora L. c. ememicus – central Arizona to Sonora L. c. festinus L. c. magdalenae L. c. martirensis L. c. melanotis – South Dakota to Iowa and central Texas L. c. merriamai – south-central and southeastern Texas to Tamaulipas L. c. richardsonii – central California L. c. sheldoni L. c. texianus – southeastern Utah and southwestern Colorado to Zacatecas L. c. wallawalla – eastern Washington to northeastern California and northwestern Nevada L. c. xanti The black-tailed jackrabbit occupies plant communities with a mixture of shrubs and forbs.
Shrubland-herb mosaics are preferred over pure stands of herbs. Black-tailed jackrabbit populations are common in sagebrush and other desert shrublands, it is common in and near croplands alfalfa fields. Ma
International Union for Conservation of Nature
The International Union for Conservation of Nature is an international organization working in the field of nature conservation and sustainable use of natural resources. It is involved in data gathering and analysis, field projects and education. IUCN's mission is to "influence and assist societies throughout the world to conserve nature and to ensure that any use of natural resources is equitable and ecologically sustainable". Over the past decades, IUCN has widened its focus beyond conservation ecology and now incorporates issues related to sustainable development in its projects. Unlike many other international environmental organisations, IUCN does not itself aim to mobilize the public in support of nature conservation, it tries to influence the actions of governments and other stakeholders by providing information and advice, through building partnerships. The organization is best known to the wider public for compiling and publishing the IUCN Red List of Threatened Species, which assesses the conservation status of species worldwide.
IUCN has a membership of over 1400 non-governmental organizations. Some 16,000 scientists and experts participate in the work of IUCN commissions on a voluntary basis, it employs 1000 full-time staff in more than 50 countries. Its headquarters are in Switzerland. IUCN has observer and consultative status at the United Nations, plays a role in the implementation of several international conventions on nature conservation and biodiversity, it was involved in establishing the World Wide Fund for Nature and the World Conservation Monitoring Centre. In the past, IUCN has been criticized for placing the interests of nature over those of indigenous peoples. In recent years, its closer relations with the business sector have caused controversy. IUCN was established in 1948, it was called the International Union for the Protection of Nature and the World Conservation Union. Establishment IUCN was established on 5 October 1948, in Fontainebleau, when representatives of governments and conservation organizations signed a formal act constituting the International Union for the Protection of Nature.
The initiative to set up the new organisation came from UNESCO and from its first Director General, the British biologist Julian Huxley. The objectives of the new Union were to encourage international cooperation in the protection of nature, to promote national and international action and to compile and distribute information. At the time of its founding IUPN was the only international organisation focusing on the entire spectrum of nature conservation Early years: 1948–1956 IUPN started out with 65 members, its secretariat was located in Brussels. Its first work program focused on saving species and habitats and applying knowledge, advancing education, promoting international agreements and promoting conservation. Providing a solid scientific base for conservation action was the heart of all activities. IUPN and UNESCO were associated, they jointly organized the 1949 Conference on Protection of Nature. In preparation for this conference a list of gravely endangered species was drawn up for the first time, a precursor of the IUCN Red List of Threatened Species.
In the early years of its existence IUCN depended entirely on UNESCO funding and was forced to temporarily scale down activities when this ended unexpectedly in 1954. IUPN was successful in engaging prominent scientists and identifying important issues such as the harmful effects of pesticides on wildlife but not many of the ideas it developed were turned into action; this was caused by unwillingness to act on the part of governments, uncertainty about the IUPN mandate and lack of resources. In 1956, IUPN changed its name to International Union for Conservation of Nature and Natural Resources. Increased profile and recognition: 1956–1965 In the 1950s and 1960s Europe entered a period of economic growth and formal colonies became independent. Both developments had impact on the work of IUCN. Through the voluntary involvement of experts in its Commissions IUCN was able to get a lot of work done while still operating on a low budget, it established links with the Council of Europe. In 1961, at the request of United Nations Economic and Social Council, the United Nations Economic and Social Council, IUCN published the first global list of national parks and protected areas which it has updated since.
IUCN's best known publication, the Red Data Book on the conservation status of species, was first published in 1964. IUCN began to play a part in the development of international treaties and conventions, starting with the African Convention on the Conservation of Nature and Natural Resources. Environmental law and policy making became a new area of expertise. Africa was the focus of many of the early IUCN conservation field projects. IUCN supported the ‘Yellowstone model’ of protected area management, which restricted human presence and activity in order to protect nature. IUCN and other conservation organisations were criticized for protecting nature against people rather than with people; this model was also applied in Africa and played a role in the decision to remove the Maasai people from Serengeti National Park and the Ngorongoro Conservation Area. To establish a stable financial basis for its work, IUCN participated in setting up the World Wildlife Fund
Pollen is a fine to coarse powdery substance comprising pollen grains which are male microgametophytes of seed plants, which produce male gametes. Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male cone to the female cone of coniferous plants. If pollen lands on a compatible pistil or female cone, it germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require magnification to see detail; the study of pollen is called palynology and is useful in paleoecology, paleontology and forensics. Pollen in plants is used for transferring haploid male genetic material from the anther of a single flower to the stigma of another in cross-pollination. In a case of self-pollination, this process takes place from the anther of a flower to the stigma of the same flower. Pollen is used as food and food supplement.
However, because of agricultural practices, it is contaminated by agricultural pesticides. Pollen itself is not the male gamete; each pollen grain contains a generative cell. In flowering plants the vegetative tube cell produces the pollen tube, the generative cell divides to form the two sperm cells. Pollen is produced in the microsporangia in the male cone of a conifer or other gymnosperm or in the anthers of an angiosperm flower. Pollen grains come in a wide variety of shapes and surface markings characteristic of the species. Pollen grains of pines and spruces are winged; the smallest pollen grain, that of the forget-me-not, is around 6 µm in diameter. Wind-borne pollen grains can be as large as about 90–100 µm. In angiosperms, during flower development the anther is composed of a mass of cells that appear undifferentiated, except for a differentiated dermis; as the flower develops, four groups of sporogenous cells form within the anther. The fertile sporogenous cells are surrounded by layers of sterile cells that grow into the wall of the pollen sac.
Some of the cells grow into nutritive cells that supply nutrition for the microspores that form by meiotic division from the sporogenous cells. In a process called microsporogenesis, four haploid microspores are produced from each diploid sporogenous cell, after meiotic division. After the formation of the four microspores, which are contained by callose walls, the development of the pollen grain walls begins; the callose wall is broken down by an enzyme called callase and the freed pollen grains grow in size and develop their characteristic shape and form a resistant outer wall called the exine and an inner wall called the intine. The exine is. Two basic types of microsporogenesis are recognised and successive. In simultaneous microsporogenesis meiotic steps I and II are completed prior to cytokinesis, whereas in successive microsporogenesis cytokinesis follows. While there may be a continuum with intermediate forms, the type of microsporogenesis has systematic significance; the predominant form amongst the monocots is successive.
During microgametogenesis, the unicellular microspores undergo mitosis and develop into mature microgametophytes containing the gametes. In some flowering plants, germination of the pollen grain may begin before it leaves the microsporangium, with the generative cell forming the two sperm cells. Except in the case of some submerged aquatic plants, the mature pollen grain has a double wall; the vegetative and generative cells are surrounded by a thin delicate wall of unaltered cellulose called the endospore or intine, a tough resistant outer cuticularized wall composed of sporopollenin called the exospore or exine. The exine bears spines or warts, or is variously sculptured, the character of the markings is of value for identifying genus, species, or cultivar or individual; the spines may be less than a micron in length referred to as spinulose, or longer than a micron referred to as echinate. Various terms describe the sculpturing such as reticulate, a net like appearance consisting of elements separated from each other by a lumen.
The pollen wall protects the sperm. The pollen grain surface is covered with waxes and proteins, which are held in place by structures called sculpture elements on the surface of the grain; the outer pollen wall, which prevents the pollen grain from shrinking and crushing the genetic material during desiccation, is composed of two layers. These two layers are the tectum and the foot layer, just above the intine; the tectum and foot layer are separated by a region called the columella, composed of strengthening rods. The outer wall is constructed with a resistant biopolymer called sporopollenin. Pollen apertures are regions of the pollen wall that may involve exine thinning or a significant reduction in exine thickness, they allow shrinking and swelling of the grain caused by changes in moisture content. Elongated apertures or furrows in the pollen grain are called sulci. Apertures that are more circular are called pores. Colpi and pores are major features in the identification of classes of pollen.
Pollen may be referre
A seed is an embryonic plant enclosed in a protective outer covering. The formation of the seed is part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosperm plants. Seeds are the product of the ripened ovule, after fertilization by pollen and some growth within the mother plant; the embryo is developed from the seed coat from the integuments of the ovule. Seeds have been an important development in the reproduction and success of gymnosperm and angiosperm plants, relative to more primitive plants such as ferns and liverworts, which do not have seeds and use water-dependent means to propagate themselves. Seed plants now dominate biological niches on land, from forests to grasslands both in hot and cold climates; the term "seed" has a general meaning that antedates the above – anything that can be sown, e.g. "seed" potatoes, "seeds" of corn or sunflower "seeds". In the case of sunflower and corn "seeds", what is sown is the seed enclosed in a shell or husk, whereas the potato is a tuber.
Many structures referred to as "seeds" are dry fruits. Plants producing berries are called baccate. Sunflower seeds are sometimes sold commercially while still enclosed within the hard wall of the fruit, which must be split open to reach the seed. Different groups of plants have other modifications, the so-called stone fruits have a hardened fruit layer fused to and surrounding the actual seed. Nuts are the one-seeded, hard-shelled fruit of some plants with an indehiscent seed, such as an acorn or hazelnut. Seeds are produced in several related groups of plants, their manner of production distinguishes the angiosperms from the gymnosperms. Angiosperm seeds are produced in a hard or fleshy structure called a fruit that encloses the seeds for protection in order to secure healthy growth; some fruits have layers of both fleshy material. In gymnosperms, no special structure develops to enclose the seeds, which begin their development "naked" on the bracts of cones. However, the seeds do become covered by the cone scales.
Seed production in natural plant populations varies from year to year in response to weather variables and diseases, internal cycles within the plants themselves. Over a 20-year period, for example, forests composed of loblolly pine and shortleaf pine produced from 0 to nearly 5 million sound pine seeds per hectare. Over this period, there were six bumper, five poor, nine good seed crops, when evaluated for production of adequate seedlings for natural forest reproduction. Angiosperm seeds consist of three genetically distinct constituents: the embryo formed from the zygote, the endosperm, triploid, the seed coat from tissue derived from the maternal tissue of the ovule. In angiosperms, the process of seed development begins with double fertilization, which involves the fusion of two male gametes with the egg cell and the central cell to form the primary endosperm and the zygote. Right after fertilization, the zygote is inactive, but the primary endosperm divides to form the endosperm tissue.
This tissue becomes the food the young plant will consume until the roots have developed after germination. After fertilization the ovules develop into the seeds; the ovule consists of a number of components: The funicle or seed stalk which attaches the ovule to the placenta and hence ovary or fruit wall, at the pericarp. The nucellus, the remnant of the megasporangium and main region of the ovule where the megagametophyte develops; the micropyle, a small pore or opening in the apex of the integument of the ovule where the pollen tube enters during the process of fertilization. The chalaza, the base of the ovule opposite the micropyle, where integument and nucellus are joined together; the shape of the ovules as they develop affects the final shape of the seeds. Plants produce ovules of four shapes: the most common shape is called anatropous, with a curved shape. Orthotropous ovules are straight with all the parts of the ovule lined up in a long row producing an uncurved seed. Campylotropous ovules have a curved megagametophyte giving the seed a tight "C" shape.
The last ovule shape is called amphitropous, where the ovule is inverted and turned back 90 degrees on its stalk. In the majority of flowering plants, the zygote's first division is transversely oriented in regards to the long axis, this establishes the polarity of the embryo; the upper or chalazal pole becomes the main area of growth of the embryo, while the lower or micropylar pole produces the stalk-like suspensor that attaches to the micropyle. The suspensor absorbs and manufactures nutrients from the endosperm that are used during the embryo's growth; the main components of the embryo are: The cotyledons, the seed leaves, attached to the embryonic axis. There may be two; the cotyledons are the source of nutrients in the non-endospermic dicotyledons, in which case they replace the endosperm, are thick and leathery. In endospermic seeds the cotyledons are papery. Dicotyledons have the point of attachment opposite one another on the axis; the epicotyl, the embryonic axis above the point of attachment of the cotyledon.
The plumule, the tip of the epicotyl, has a feathery appearance due to the presence of young leaf primordia at the apex, will become the shoot upon germination. The hypocotyl, the embryonic axis below the point of attachment of the cotyledon, connecting the epicotyl and the radicle, being the stem-root transition zone; the radicle, the basal tip of the hy
Nevada is a state in the Western United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast and Utah to the east. Nevada is the 7th most extensive, the 32nd most populous, but the 9th least densely populated of the U. S. states. Nearly three-quarters of Nevada's people live in Clark County, which contains the Las Vegas–Paradise metropolitan area where three of the state's four largest incorporated cities are located. Nevada's capital, however, is Carson City. Nevada is known as the "Silver State" because of the importance of silver to its history and economy, it is known as the "Battle Born State", because it achieved statehood during the Civil War. Nevada is desert and semi-arid, much of it within the Great Basin. Areas south of the Great Basin are within the Mojave Desert, while Lake Tahoe and the Sierra Nevada lie on the western edge. About 86% of the state's land is managed by various jurisdictions of the U. S. federal government, both civilian and military.
Before European contact, Native Americans of the Paiute and Washoe tribes inhabited the land, now Nevada. The first Europeans to explore the region were Spanish, they called the region Nevada because of the snow. The area formed part of the Viceroyalty of New Spain, became part of Mexico when it gained independence in 1821; the United States annexed the area in 1848 after its victory in the Mexican–American War, it was incorporated as part of Utah Territory in 1850. The discovery of silver at the Comstock Lode in 1859 led to a population boom that became an impetus to the creation of Nevada Territory out of western Utah Territory in 1861. Nevada became the 36th state on October 31, 1864, as the second of two states added to the Union during the Civil War. Nevada has a reputation for its libertarian laws. In 1940, with a population of just over 110,000 people, Nevada was by far the least-populated state, with less than half the population of the next least-populated state. However, legalized gambling and lenient marriage and divorce laws transformed Nevada into a major tourist destination in the 20th century.
Nevada is the only U. S. state where prostitution is legal, though it is illegal in Clark County, Washoe County and Carson City. The tourism industry remains Nevada's largest employer, with mining continuing as a substantial sector of the economy: Nevada is the fourth-largest producer of gold in the world; the name "Nevada" comes from meaning "snow-covered", after the Sierra Nevada. Most Nevadans pronounce the second syllable of their state name using the TRAP vowel. Many from outside the Western United States pronounce it with the PALM vowel. Although the latter pronunciation is closer to the Spanish pronunciation, it is not the pronunciation preferred by most Nevadans. State Assemblyman Harry Mortenson proposed a bill to recognize the alternate pronunciation of Nevada, though the bill was not supported by most legislators and never received a vote; the Nevadan pronunciation is the de facto official one, since it is the one used by the state legislature. At one time, the state's official tourism organization, TravelNevada, stylized the name of the state as "Nevăda", with a breve mark over the a indicating the locally preferred pronunciation, available as a license plate design.
Nevada is entirely within the Basin and Range Province, is broken up by many north-south mountain ranges. Most of these ranges have endorheic valleys between them, which belies the image portrayed by the term Great Basin. Much of the northern part of the state is within the Great Basin, a mild desert that experiences hot temperatures in the summer and cold temperatures in the winter. Moisture from the Arizona Monsoon will cause summer thunderstorms; the state's highest recorded temperature was 125 °F in Laughlin on June 29, 1994. The coldest recorded temperature was −52 °F set in San Jacinto in 1972, in the northeastern portion of the state; the Humboldt River crosses the state from east to west across the northern part of the state, draining into the Humboldt Sink near Lovelock. Several rivers drain from the Sierra Nevada eastward, including the Walker and Carson rivers. All of these rivers are endorheic basins, ending in Walker Lake, Pyramid Lake, the Carson Sink, respectively. However, not all of Nevada is within the Great Basin.
Tributaries of the Snake River drain the far north, while the Colorado River, which forms much of the boundary with Arizona, drains much of southern Nevada. The mountain ranges, some of which have peaks above 13,000 feet, harbor lush forests high above desert plains, creating sky islands for endemic species; the valleys are no lower in elevation than 3,000 feet, while some in central Nevada are above 6,000 feet. The southern third of the state, where the Las Vegas area is situated, is within the Mojave Desert; the area is closer to the Arizona Monsoon in the summer. The terrain is lower below 4,000 feet, creating conditions for hot summer days and cool to chilly winter nights. Nevada and California have by far the longest diagonal line as a state boundary at just over 400 miles; this line begins in Lake Tahoe nearly
A leaf is an organ of a vascular plant and is the principal lateral appendage of the stem. The leaves and stem together form the shoot. Leaves are collectively referred to as foliage, as in "autumn foliage". A leaf is a thin, dorsiventrally flattened organ borne above ground and specialized for photosynthesis. In most leaves, the primary photosynthetic tissue, the palisade mesophyll, is located on the upper side of the blade or lamina of the leaf but in some species, including the mature foliage of Eucalyptus, palisade mesophyll is present on both sides and the leaves are said to be isobilateral. Most leaves have distinct upper surface and lower surface that differ in colour, the number of stomata, the amount and structure of epicuticular wax and other features. Leaves can have many different shapes and textures; the broad, flat leaves with complex venation of flowering plants are known as megaphylls and the species that bear them, the majority, as broad-leaved or megaphyllous plants. In the clubmosses, with different evolutionary origins, the leaves are simple and are known as microphylls.
Some leaves, such as bulb scales, are not above ground. In many aquatic species the leaves are submerged in water. Succulent plants have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls and spines. Furthermore, several kinds of leaf-like structures found in vascular plants are not homologous with them. Examples include flattened plant stems called phylloclades and cladodes, flattened leaf stems called phyllodes which differ from leaves both in their structure and origin; some structures of non-vascular plants function much like leaves. Examples include the phyllids of liverworts. Leaves are the most important organs of most vascular plants. Green plants are autotrophic, meaning that they do not obtain food from other living things but instead create their own food by photosynthesis, they capture the energy in sunlight and use it to make simple sugars, such as glucose and sucrose, from carbon dioxide and water. The sugars are stored as starch, further processed by chemical synthesis into more complex organic molecules such as proteins or cellulose, the basic structural material in plant cell walls, or metabolised by cellular respiration to provide chemical energy to run cellular processes.
The leaves draw water from the ground in the transpiration stream through a vascular conducting system known as xylem and obtain carbon dioxide from the atmosphere by diffusion through openings called stomata in the outer covering layer of the leaf, while leaves are orientated to maximise their exposure to sunlight. Once sugar has been synthesized, it needs to be transported to areas of active growth such as the plant shoots and roots. Vascular plants transport sucrose in a special tissue called the phloem; the phloem and xylem are parallel to each other but the transport of materials is in opposite directions. Within the leaf these vascular systems branch to form veins which supply as much of the leaf as possible, ensuring that cells carrying out photosynthesis are close to the transportation system. Leaves are broad and thin, thereby maximising the surface area directly exposed to light and enabling the light to penetrate the tissues and reach the chloroplasts, thus promoting photosynthesis.
They are arranged on the plant so as to expose their surfaces to light as efficiently as possible without shading each other, but there are many exceptions and complications. For instance plants adapted to windy conditions may have pendent leaves, such as in many willows and eucalyptss; the flat, or laminar, shape maximises thermal contact with the surrounding air, promoting cooling. Functionally, in addition to carrying out photosynthesis, the leaf is the principal site of transpiration, providing the energy required to draw the transpiration stream up from the roots, guttation. Many gymnosperms have thin needle-like or scale-like leaves that can be advantageous in cold climates with frequent snow and frost; these are interpreted as reduced from megaphyllous leaves of their Devonian ancestors. Some leaf forms are adapted to modulate the amount of light they absorb to avoid or mitigate excessive heat, ultraviolet damage, or desiccation, or to sacrifice light-absorption efficiency in favour of protection from herbivory.
For xerophytes the major constraint drought. Some window plants such as Fenestraria species and some Haworthia species such as Haworthia tesselata and Haworthia truncata are examples of xerophytes. and Bulbine mesembryanthemoides. Leaves function to store chemical energy and water and may become specialised organs serving other functions, such as tendrils of peas and other legumes, the protective spines of cacti and the insect traps in carnivorous plants such as Nepenthes and Sarracenia. Leaves are the fundamental structural units from which cones are constructed in gymnosperms and from which flowers are constructed in flowering plants; the internal organisation of most kinds of leaves has evolved to maximise exposure of the photosynthetic organelles, the chloroplasts, to light and to increase the absorption of carbon dioxide while at the same time controlling water loss. Their surfaces are waterproofed by the plant cuticle and gas exchange between the mesophyll cells and the atmosphere is controlled by minute openings called stomata which open or close to regulate the rate exchange of carbon dioxide and water vapour into
Wyoming is a state in the mountain region of the western United States. The state is the 10th largest by area, the least populous, the second most sparsely populated state in the country. Wyoming is bordered on the north by Montana, on the east by South Dakota and Nebraska, on the south by Colorado, on the southwest by Utah, on the west by Idaho and Montana; the state population was estimated at 577,737 in 2018, less than 31 of the most populous U. S. cities including Denver in neighboring Colorado. Cheyenne is the state capital and the most populous city, with an estimated population of 63,624 in 2017; the western two-thirds of the state is covered by the mountain ranges and rangelands of the Rocky Mountains, while the eastern third of the state is high elevation prairie called the High Plains. Half of the land in Wyoming is owned by the U. S. government, leading Wyoming to rank sixth by area and fifth by proportion of a state's land owned by the federal government. Federal lands include two national parks—Grand Teton and Yellowstone—two national recreation areas, two national monuments, several national forests, historic sites, fish hatcheries, wildlife refuges.
Original inhabitants of the region include the Crow, Arapaho and Shoshone. Southwestern Wyoming was in the Spanish Empire and Mexican territory until it was ceded to the United States in 1848 at the end of the Mexican–American War; the region acquired the name Wyoming when a bill was introduced to the U. S. Congress in 1865 to provide a "temporary government for the territory of Wyoming"; the name was used earlier for the Wyoming Valley in Pennsylvania, is derived from the Munsee word xwé:wamənk, meaning "at the big river flat". The main drivers of Wyoming's economy are mineral extraction—mostly coal, natural gas, trona—and tourism. Agricultural commodities include livestock, sugar beets and wool; the climate is semi-arid and continental and windier than the rest of the U. S. with greater temperature extremes. Wyoming has been a politically conservative state since the 1950s, with the Republican Party candidate winning every presidential election except 1964. Wyoming's climate is semi-arid and continental, is drier and windier in comparison to most of the United States with greater temperature extremes.
Much of this is due to the topography of the state. Summers in Wyoming are warm with July high temperatures averaging between 85 and 95 °F in most of the state. With increasing elevation, this average drops with locations above 9,000 feet averaging around 70 °F. Summer nights throughout the state are characterized by a rapid cooldown with the hottest locations averaging in the 50–60 °F range at night. In most of the state, most of the precipitation tends to fall in early summer. Winters are cold, but are variable with periods of sometimes extreme cold interspersed between mild periods, with Chinook winds providing unusually warm temperatures in some locations. Wyoming is a dry state with much of the land receiving less than 10 inches of rainfall per year. Precipitation depends on elevation with lower areas in the Big Horn Basin averaging 5–8 inches; the lower areas in the North and on the eastern plains average around 10–12 inches, making the climate there semi-arid. Some mountain areas do receive a good amount of precipitation, 20 inches or more, much of it as snow, sometimes 200 inches or more annually.
The state's highest recorded temperature is 114 °F at Basin on July 12, 1900 and the lowest recorded temperature is −66 °F at Riverside on February 9, 1933. The number of thunderstorm days vary across the state with the southeastern plains of the state having the most days of thunderstorm activity. Thunderstorm activity in the state is highest during early summer; the southeastern corner of the state is the most vulnerable part of the state to tornado activity. Moving away from that point and westwards, the incidence of tornadoes drops with the west part of the state showing little vulnerability. Tornadoes, where they occur, tend to be small and brief, unlike some of those that occur farther east; as specified in the designating legislation for the Territory of Wyoming, Wyoming's borders are lines of latitude 41°N and 45°N, longitude 104°3'W and 111°3'W, making the shape of the state a latitude-longitude quadrangle. Wyoming is one of only three states to have borders along only straight latitudinal and longitudinal lines, rather than being defined by natural landmarks.
Due to surveying inaccuracies during the 19th century, Wyoming's legal border deviates from the true latitude and longitude lines by up to half of a mile in some spots in the mountainous region along the 45th parallel. Wyoming is bordered on the north by Montana, on the east by South Dakota and Nebraska, on the south by Colorado, on the southwest by Utah, on the west by Idaho, it is the tenth largest state in the United States in total area, containing 97,814 square miles and is made up of 23 counties. From the north border to the south border it is 276 miles; the Great Plains meet the Rocky Mountains in Wyoming. The state is a great plateau broken by many mountain ranges. Surface elevations range from the summit of Gannett Peak in the Wind River Mountain Range, at 13,804 feet, to the Belle Fourche River val