Suriname known as the Republic of Suriname, is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north, French Guiana to the east, Guyana to the west and Brazil to the south. At just under 165,000 square kilometers, it is the smallest sovereign state in South America. Suriname has a population of 558,368, most of whom live on the country's north coast, in and around the capital and largest city, Paramaribo. Suriname was long inhabited by various indigenous people before being invaded and contested by European powers from the 16th century coming under Dutch rule in the late 17th century; as the chief sugar colony during the Dutch colonial period, it was a plantation economy dependent on African slaves and, following the abolition of slavery in 1863, indentured servants from Asia. Suriname was ruled by the Dutch-chartered company Sociëteit van Suriname between 1683 and 1795. In 1954, Suriname became one of the constituent countries of the Kingdom of the Netherlands.
On 25 November 1975, the country of Suriname left the Kingdom of the Netherlands to become an independent state, nonetheless maintaining close economic and cultural ties to its former colonizer. Suriname is considered to be a culturally Caribbean country, is a member of the Caribbean Community. While Dutch is the official language of government, business and education, Sranan Tongo, an English-based creole language, is a used lingua franca. Suriname is the only sovereign nation outside Europe where Dutch is spoken by a majority of the population; as a legacy of colonization, the people of Suriname are among the most diverse in the world, spanning a multitude of ethnic and linguistic groups. The name Suriname may derive from an indigenous people called Surinen, who inhabited the area at the time of European contact. British settlers, who founded the first European colony at Marshall's Creek along the Suriname River, spelled the name as "Surinam"; when the territory was taken over by the Dutch, it became part of a group of colonies known as Dutch Guiana.
The official spelling of the country's English name was changed from "Surinam" to "Suriname" in January 1978, but "Surinam" can still be found in English. A notable example is Surinam Airways; the older English name is reflected in the English pronunciation. In Dutch, the official language of Suriname, the pronunciation is, with the main stress on the third syllable and a schwa terminal vowel. Indigenous settlement of Suriname dates back to 3,000 BC; the largest tribes were a nomadic coastal tribe that lived from hunting and fishing. They were the first inhabitants in the area; the Carib settled in the area and conquered the Arawak by using their superior sailing ships. They settled in Galibi at the mouth of the Marowijne River. While the larger Arawak and Carib tribes lived along the coast and savanna, smaller groups of indigenous people lived in the inland rainforest, such as the Akurio, Trió, Wayana. Beginning in the 16th century, French and English explorers visited the area. A century Dutch and English settlers established plantation colonies along the many rivers in the fertile Guiana plains.
The earliest documented colony in Guiana was an English settlement named Marshall's Creek along the Suriname River. After that there was another short-lived English colony called Willoughbyland that lasted from 1650 to 1674. Disputes arose between the English for control of this territory. In 1667, during negotiations leading to the Treaty of Breda, the Dutch decided to keep the nascent plantation colony of Suriname they had gained from the English; the English were able to keep New Amsterdam, the main city of the former colony of New Netherland in North America on the mid-Atlantic coast. A cultural and economic hub in those days, they renamed it after the Duke of York: New York City. In 1683, the Society of Suriname was founded by the city of Amsterdam, the Van Aerssen van Sommelsdijck family, the Dutch West India Company; the society was chartered to defend the colony. The planters of the colony relied on African slaves to cultivate and process the commodity crops of coffee, sugar cane and cotton plantations along the rivers.
Planters' treatment of the slaves was notoriously bad—historian C. R. Boxer wrote that "man's inhumanity to man just about reached its limits in Surinam"—and many slaves escaped the plantations. With the help of the native South Americans living in the adjoining rain forests, these runaway slaves established a new and unique culture in the interior, successful in its own right, they were known collectively in English as Maroons, in French as Nèg'Marrons, in Dutch as Marrons. The Maroons developed several independent tribes through a process of ethnogenesis, as they were made up of slaves from different African ethnicities; these tribes include the Saramaka, Ndyuka or Aukan, Aluku or Boni, Matawai. The Maroons raided plantations to recruit new members from the slaves and capture women, as well as to acquire weapons and supplies, they sometimes killed their families in the raids. The colonists mounted armed campaigns against the Maroons, who escaped through the rain forest, which they knew much better than did the colonis
The Holocene is the current geological epoch. It began 11,650 cal years before present, after the last glacial period, which concluded with the Holocene glacial retreat; the Holocene and the preceding Pleistocene together form the Quaternary period. The Holocene has been identified with the current warm period, known as MIS 1, it is considered by some to be an interglacial period within the Pleistocene Epoch. The Holocene has seen the growth and impacts of the human species worldwide, including all its written history, development of major civilizations, overall significant transition toward urban living in the present. Human impacts on modern-era Earth and its ecosystems may be considered of global significance for future evolution of living species, including synchronous lithospheric evidence, or more hydrospheric and atmospheric evidence of human impacts. In July 2018, the International Union of Geological Sciences split the Holocene epoch into three distinct subsections, Greenlandian and Meghalayan, as proposed by International Commission on Stratigraphy.
The boundary stratotype of Meghalayan is a speleothem in Mawmluh cave in India, the global auxiliary stratotype is an ice core from Mount Logan in Canada. The name Holocene comes from the Ancient Greek words ὅλος and καινός, meaning "entirely recent", it is accepted by the International Commission on Stratigraphy that the Holocene started 11,650 cal years BP. The Subcommission on Quaternary Stratigraphy quotes Gibbard and van Kolfschoten in Gradstein Ogg and Smith in stating the term'Recent' as an alternative to Holocene is invalid and should not be used and observe that the term Flandrian, derived from marine transgression sediments on the Flanders coast of Belgium has been used as a synonym for Holocene by authors who consider the last 10,000 years should have the same stage-status as previous interglacial events and thus be included in the Pleistocene; the International Commission on Stratigraphy, considers the Holocene an epoch following the Pleistocene and the last glacial period. Local names for the last glacial period include the Wisconsinan in North America, the Weichselian in Europe, the Devensian in Britain, the Llanquihue in Chile and the Otiran in New Zealand.
The Holocene can be subdivided into five time intervals, or chronozones, based on climatic fluctuations: Preboreal, Atlantic and Subatlantic. Note: "ka" means "kilo-annum" Before Present, i.e. 1,000 years before 1950 The Blytt–Sernander classification of climatic periods defined by plant remains in peat mosses, is being explored. Geologists working in different regions are studying sea levels, peat bogs and ice core samples by a variety of methods, with a view toward further verifying and refining the Blytt–Sernander sequence, they find a general correspondence across Eurasia and North America, though the method was once thought to be of no interest. The scheme was defined for Northern Europe, but the climate changes were claimed to occur more widely; the periods of the scheme include a few of the final pre-Holocene oscillations of the last glacial period and classify climates of more recent prehistory. Paleontologists have not defined any faunal stages for the Holocene. If subdivision is necessary, periods of human technological development, such as the Mesolithic and Bronze Age, are used.
However, the time periods referenced by these terms vary with the emergence of those technologies in different parts of the world. Climatically, the Holocene may be divided evenly into the Neoglacial periods. According to some scholars, a third division, the Anthropocene, has now begun; the International Commission on Stratigraphy Subcommission on Quaternary Stratigraphy’s working group on the'Anthropocene' note this term is used to denote the present time interval in which many geologically significant conditions and processes have been profoundly altered by human activities. The'Anthropocene' is not a formally defined geological unit. Continental motions due to plate tectonics are less than a kilometre over a span of only 10,000 years. However, ice melt caused world sea levels to rise about 35 m in the early part of the Holocene. In addition, many areas above about 40 degrees north latitude had been depressed by the weight of the Pleistocene glaciers and rose as much as 180 m due to post-glacial rebound over the late Pleistocene and Holocene, are still rising today.
The sea level rise and temporary land depression allowed temporary marine incursions into areas that are now far from the sea. Holocene marine fossils are known, from Vermont and Michigan. Other than higher-latitude temporary marine incursions associated with glacial depression, Holocene fossils are found in lakebed and cave deposits. Holocene marine deposits along low-latitude coastlines are rare because the rise in sea levels during the period exceeds any tectonic uplift of non-glacial origin. Post-glacial rebound in the Scandinavia region resulted in the formation of the Baltic Sea; the region continues to rise, still causing weak earthquakes across Northern Europe. The equivalent event in North America was the rebound of Hudson Bay, as it shrank from its larger, immediate post-glacial Tyrrell Sea phase, to near its present boundaries. Climate has been stable over the Holocene. Ice core
The Cretaceous is a geologic period and system that spans 79 million years from the end of the Jurassic Period 145 million years ago to the beginning of the Paleogene Period 66 mya. It is the last period of the Mesozoic Era, the longest period of the Phanerozoic Eon; the Cretaceous Period is abbreviated K, for its German translation Kreide. The Cretaceous was a period with a warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas; these oceans and seas were populated with now-extinct marine reptiles and rudists, while dinosaurs continued to dominate on land. During this time, new groups of mammals and birds, as well as flowering plants, appeared; the Cretaceous ended with the Cretaceous–Paleogene extinction event, a large mass extinction in which many groups, including non-avian dinosaurs and large marine reptiles died out. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary, a geologic signature associated with the mass extinction which lies between the Mesozoic and Cenozoic eras.
The Cretaceous as a separate period was first defined by Belgian geologist Jean d'Omalius d'Halloy in 1822, using strata in the Paris Basin and named for the extensive beds of chalk, found in the upper Cretaceous of Western Europe. The name Cretaceous was derived from Latin creta; the Cretaceous is divided into Early and Late Cretaceous epochs, or Lower and Upper Cretaceous series. In older literature the Cretaceous is sometimes divided into three series: Neocomian and Senonian. A subdivision in eleven stages, all originating from European stratigraphy, is now used worldwide. In many parts of the world, alternative local subdivisions are still in use; as with other older geologic periods, the rock beds of the Cretaceous are well identified but the exact age of the system's base is uncertain by a few million years. No great extinction or burst of diversity separates the Cretaceous from the Jurassic. However, the top of the system is defined, being placed at an iridium-rich layer found worldwide, believed to be associated with the Chicxulub impact crater, with its boundaries circumscribing parts of the Yucatán Peninsula and into the Gulf of Mexico.
This layer has been dated at 66.043 Ma. A 140 Ma age for the Jurassic-Cretaceous boundary instead of the accepted 145 Ma was proposed in 2014 based on a stratigraphic study of Vaca Muerta Formation in Neuquén Basin, Argentina. Víctor Ramos, one of the authors of the study proposing the 140 Ma boundary age sees the study as a "first step" toward formally changing the age in the International Union of Geological Sciences. From youngest to oldest, the subdivisions of the Cretaceous period are: Late Cretaceous Maastrichtian – Campanian – Santonian – Coniacian – Turonian – Cenomanian – Early Cretaceous Albian – Aptian – Barremian – Hauterivian – Valanginian – Berriasian – The high sea level and warm climate of the Cretaceous meant large areas of the continents were covered by warm, shallow seas, providing habitat for many marine organisms; the Cretaceous was named for the extensive chalk deposits of this age in Europe, but in many parts of the world, the deposits from the Cretaceous are of marine limestone, a rock type, formed under warm, shallow marine circumstances.
Due to the high sea level, there was extensive space for such sedimentation. Because of the young age and great thickness of the system, Cretaceous rocks are evident in many areas worldwide. Chalk is a rock type characteristic for the Cretaceous, it consists of coccoliths, microscopically small calcite skeletons of coccolithophores, a type of algae that prospered in the Cretaceous seas. In northwestern Europe, chalk deposits from the Upper Cretaceous are characteristic for the Chalk Group, which forms the white cliffs of Dover on the south coast of England and similar cliffs on the French Normandian coast; the group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not consolidated and the Chalk Group still consists of loose sediments in many places; the group has other limestones and arenites. Among the fossils it contains are sea urchins, belemnites and sea reptiles such as Mosasaurus. In southern Europe, the Cretaceous is a marine system consisting of competent limestone beds or incompetent marls.
Because the Alpine mountain chains did not yet exist in the Cretaceous, these deposits formed on the southern edge of the European continental shelf, at the margin of the Tethys Ocean. Stagnation of deep sea currents in middle Cretaceous times caused anoxic conditions in the sea water leaving the deposited organic matter undecomposed. Half the worlds petroleum reserves were laid down at this time in the anoxic conditions of what would become the Persian Gulf and the Gulf of Mexico. In many places around the world, dark anoxic shales were formed during this interval; these shales are an important source rock for oil and gas, for example in the subsurface of the North Sea. During th
In vascular plants, the root is the organ of a plant that lies below the surface of the soil. Roots can be aerial or aerating, that is, growing up above the ground or above water. Furthermore, a stem occurring below ground is not exceptional either. Therefore, the root is best defined as the non-nodes bearing parts of the plant's body. However, important internal structural differences between stems and roots exist; the fossil record of roots—or rather, infilled voids where roots rotted after death—spans back to the late Silurian, about 430 million years ago. Their identification is difficult, because casts and molds of roots are so similar in appearance to animal burrows, they can be discriminated using a range of features. The first root that comes from a plant is called the radicle. A root's four major functions are: absorption of inorganic nutrients. In response to the concentration of nutrients, roots synthesise cytokinin, which acts as a signal as to how fast the shoots can grow. Roots function in storage of food and nutrients.
The roots of most vascular plant species enter into symbiosis with certain fungi to form mycorrhizae, a large range of other organisms including bacteria closely associate with roots. When dissected, the arrangement of the cells in a root is root hair, epiblem, endodermis, pericycle and, the vascular tissue in the centre of a root to transport the water absorbed by the root to other places of the plant; the most striking characteristic of roots is that, roots have an endogenous origin, i.e. it originates and develops from an inner layer of the mother axis. Whereas Stem-branching and leaves are exogenous, i.e. start to develop from the cortex, an outer layer. In its simplest form, the term root architecture refers to the spatial configuration of a plant’s root system; this system can be complex and is dependent upon multiple factors such as the species of the plant itself, the composition of the soil and the availability of nutrients. The configuration of root systems serves to structurally support the plant, compete with other plants and for uptake of nutrients from the soil.
Roots grow to specific conditions. For example, a root system that has developed in dry soil may not be as efficient in flooded soil, yet plants are able to adapt to other changes in the environment, such as seasonal changes. Root architecture plays the important role of providing a secure supply of nutrients and water as well as anchorage and support; the main terms used to classify the architecture of a root system are: Branch magnitude: the number of links. Topology: the pattern of branching, including:Herringbone: alternate lateral branching off a parent root Dichotomous: opposite, forked branches Radial: whorl of branches around a rootLink length: the distance between branches. Root angle: the radial angle of a lateral root’s base around the parent root’s circumference, the angle of a lateral root from its parent root, the angle an entire system spreads. Link radius: the diameter of a root. All components of the root architecture are regulated through a complex interaction between genetic responses and responses due to environmental stimuli.
These developmental stimuli are categorised as intrinsic, the genetic and nutritional influences, or extrinsic, the environmental influences and are interpreted by signal transduction pathways. The extrinsic factors that affect root architecture include gravity, light exposure and oxygen, as well as the availability or lack of nitrogen, sulphur and sodium chloride; the main hormones and respective pathways responsible for root architecture development include: Auxin – Auxin promotes root initiation, root emergence and primary root elongation. Cytokinins – Cytokinins regulate root apical meristem size and promote lateral root elongation. Gibberellins -- Together with ethylene they promote elongation. Together with auxin they promote root elongation. Gibberellins inhibit lateral root primordia initiation. Ethylene – Ethylene promotes crown root formation. Early root growth is one of the functions of the apical meristem located near the tip of the root; the meristem cells more or less continuously divide, producing more meristem, root cap cells, undifferentiated root cells.
The latter become the primary tissues of the root, first undergoing elongation, a process that pushes the root tip forward in the growing medium. These cells differentiate and mature into specialized cells of the root tissues. Growth from apical meristems is known as primary growth. Secondary growth encompasses all growth in diameter, a major component of woody plant tissues and many nonwoody plants. For example, storage roots of sweet potato are not woody. Secondary growth occurs at the lateral meristems, namely the vascular cork cambium; the former forms secondary phloem, while the latter forms the periderm. In plants with secondary growth, the vascular cambium, originating between the xylem and the phloem, forms a cylinder of tissue along the stem and root; the vascular cambium forms new cells on both the inside and outside of the cambium cylinder, with those on the inside forming secondary xylem cells, those on the outside forming secondary phloem cells. As
The Carboniferous is a geologic period and system that spans 60 million years from the end of the Devonian Period 358.9 million years ago, to the beginning of the Permian Period, 298.9 Mya. The name Carboniferous means "coal-bearing" and derives from the Latin words carbō and ferō, was coined by geologists William Conybeare and William Phillips in 1822. Based on a study of the British rock succession, it was the first of the modern'system' names to be employed, reflects the fact that many coal beds were formed globally during that time; the Carboniferous is treated in North America as two geological periods, the earlier Mississippian and the Pennsylvanian. Terrestrial animal life was well established by the Carboniferous period. Amphibians were the dominant land vertebrates, of which one branch would evolve into amniotes, the first terrestrial vertebrates. Arthropods were very common, many were much larger than those of today. Vast swaths of forest covered the land, which would be laid down and become the coal beds characteristic of the Carboniferous stratigraphy evident today.
The atmospheric content of oxygen reached its highest levels in geological history during the period, 35% compared with 21% today, allowing terrestrial invertebrates to evolve to great size. The half of the period experienced glaciations, low sea level, mountain building as the continents collided to form Pangaea. A minor marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred at the end of the period, caused by climate change. In the United States the Carboniferous is broken into Mississippian and Pennsylvanian subperiods; the Mississippian is about twice as long as the Pennsylvanian, but due to the large thickness of coal-bearing deposits with Pennsylvanian ages in Europe and North America, the two subperiods were long thought to have been more or less equal in duration. In Europe the Lower Carboniferous sub-system is known as the Dinantian, comprising the Tournaisian and Visean Series, dated at 362.5-332.9 Ma, the Upper Carboniferous sub-system is known as the Silesian, comprising the Namurian and Stephanian Series, dated at 332.9-298.9 Ma.
The Silesian is contemporaneous with the late Mississippian Serpukhovian plus the Pennsylvanian. In Britain the Dinantian is traditionally known as the Carboniferous Limestone, the Namurian as the Millstone Grit, the Westphalian as the Coal Measures and Pennant Sandstone; the International Commission on Stratigraphy faunal stages from youngest to oldest, together with some of their regional subdivisions, are: A global drop in sea level at the end of the Devonian reversed early in the Carboniferous. There was a drop in south polar temperatures; these conditions had little effect in the deep tropics, where lush swamps to become coal, flourished to within 30 degrees of the northernmost glaciers. Mid-Carboniferous, a drop in sea level precipitated a major marine extinction, one that hit crinoids and ammonites hard; this sea level drop and the associated unconformity in North America separate the Mississippian subperiod from the Pennsylvanian subperiod. This happened about 323 million years ago, at the onset of the Permo-Carboniferous Glaciation.
The Carboniferous was a time of active mountain-building as the supercontinent Pangaea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America–Europe along the present line of eastern North America; this continental collision resulted in the Hercynian orogeny in Europe, the Alleghenian orogeny in North America. In the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural Mountains. Most of the Mesozoic supercontinent of Pangea was now assembled, although North China, South China continents were still separated from Laurasia; the Late Carboniferous Pangaea was shaped like an "O." There were two major oceans in the Carboniferous—Panthalassa and Paleo-Tethys, inside the "O" in the Carboniferous Pangaea. Other minor oceans were shrinking and closed - Rheic Ocean, the small, shallow Ural Ocean and Proto-Tethys Ocean. Average global temperatures in the Early Carboniferous Period were high: 20 °C.
However, cooling during the Middle Carboniferous reduced average global temperatures to about 12 °C. Lack of growth rings of fossilized trees suggest a lack of seasons of a tropical climate. Glaciations in Gondwana, triggered by Gondwana's southward movement, continued into the Permian and because of the lack of clear markers and breaks, the deposits of this glacial period are referred to as Permo-Carboniferous in age; the cooling and drying of the climate led to the Carboniferous Rainforest Collapse during the late Carboniferous. Tropical rainforests fragmented and were devastated by climate change. Carboniferous rocks in Europe and eastern North America consist of a repeated sequence of limestone, sandstone and coal beds. In North America, the early Carboniferous is marine
A snout is the protruding portion of an animal's face, consisting of its nose and jaw. In many animals, the equivalent structure is called rostrum, or proboscis; the wet furless surface around the nostrils of the nose of some animals is called the rhinarium. The rhinarium is associated with a stronger sense of olfaction; the snout is considered a weak point on most animals: because of its structure, an animal can be stunned or knocked out, or have its snout snapped by applying sufficient force. Snouts are found on many mammals in a variety of shapes; some animals, including ursines and great cats, have box-like snouts, while others, like shrews, have pointed snouts. Pig snouts are cylindrical; the muzzle begins at the stop, just below the eyes, includes the dog's nose and mouth. In the domestic dog, most of the upper muzzle contains organs for detecting scents; the loose flaps of skin on the sides of the upper muzzle that hang to different lengths over the mouth are called flews. It is innervated by one of the twelve pairs of cranial nerves.
These nerves emerge through the skull to their target organs. Other destinations of these nerves are eyeballs and tongue; the muzzle shape of a domestic dog ranges in shape depending upon the breed, from long and thin, as in the Rough Collie, to nearly nonexistent because it is so flat, as in the Pug. Some breeds, such as many sled dogs and Spitz types, have muzzles that somewhat resemble the original wolf's in size and shape, others in the less extreme range have shortened it somewhat as in many hounds
In botany, a fruit is the seed-bearing structure in flowering plants formed from the ovary after flowering. Fruits are the means. Edible fruits, in particular, have propagated with the movements of humans and animals in a symbiotic relationship as a means for seed dispersal and nutrition. Accordingly, fruits account for a substantial fraction of the world's agricultural output, some have acquired extensive cultural and symbolic meanings. In common language usage, "fruit" means the fleshy seed-associated structures of a plant that are sweet or sour, edible in the raw state, such as apples, grapes, lemons and strawberries. On the other hand, in botanical usage, "fruit" includes many structures that are not called "fruits", such as bean pods, corn kernels and wheat grains; the section of a fungus that produces spores is called a fruiting body. Many common terms for seeds and fruit do not correspond to the botanical classifications. In culinary terminology, a fruit is any sweet-tasting plant part a botanical fruit.
However, in botany, a fruit is the ripened ovary or carpel that contains seeds, a nut is a type of fruit and not a seed, a seed is a ripened ovule. Examples of culinary "vegetables" and nuts that are botanically fruit include corn, eggplant, sweet pepper, tomato. In addition, some spices, such as allspice and chili pepper, are fruits. In contrast, rhubarb is referred to as a fruit, because it is used to make sweet desserts such as pies, though only the petiole of the rhubarb plant is edible, edible gymnosperm seeds are given fruit names, e.g. ginkgo nuts and pine nuts. Botanically, a cereal grain, such as corn, rice, or wheat, is a kind of fruit, termed a caryopsis. However, the fruit wall is thin and is fused to the seed coat, so all of the edible grain is a seed; the outer edible layer, is the pericarp, formed from the ovary and surrounding the seeds, although in some species other tissues contribute to or form the edible portion. The pericarp may be described in three layers from outer to inner, the epicarp and endocarp.
Fruit that bears a prominent pointed terminal projection is said to be beaked. A fruit results from maturation of one or more flowers, the gynoecium of the flower forms all or part of the fruit. Inside the ovary/ovaries are one or more ovules where the megagametophyte contains the egg cell. After double fertilization, these ovules will become seeds; the ovules are fertilized in a process that starts with pollination, which involves the movement of pollen from the stamens to the stigma of flowers. After pollination, a tube grows from the pollen through the stigma into the ovary to the ovule and two sperm are transferred from the pollen to the megagametophyte. Within the megagametophyte one of the two sperm unites with the egg, forming a zygote, the second sperm enters the central cell forming the endosperm mother cell, which completes the double fertilization process; the zygote will give rise to the embryo of the seed, the endosperm mother cell will give rise to endosperm, a nutritive tissue used by the embryo.
As the ovules develop into seeds, the ovary begins to ripen and the ovary wall, the pericarp, may become fleshy, or form a hard outer covering. In some multiseeded fruits, the extent to which the flesh develops is proportional to the number of fertilized ovules; the pericarp is differentiated into two or three distinct layers called the exocarp and endocarp. In some fruits simple fruits derived from an inferior ovary, other parts of the flower, fuse with the ovary and ripen with it. In other cases, the sepals, petals and/or stamens and style of the flower fall off; when such other floral parts are a significant part of the fruit, it is called an accessory fruit. Since other parts of the flower may contribute to the structure of the fruit, it is important to study flower structure to understand how a particular fruit forms. There are three general modes of fruit development: Apocarpous fruits develop from a single flower having one or more separate carpels, they are the simplest fruits. Syncarpous fruits develop from a single gynoecium having two or more carpels fused together.
Multiple fruits form from many different flowers. Plant scientists have grouped fruits into three main groups, simple fruits, aggregate fruits, composite or multiple fruits; the groupings are not evolutionarily relevant, since many diverse plant taxa may be in the same group, but reflect how the flower organs are arranged and how the fruits develop. Simple fruits can be either dry or fleshy, result from the ripening of a simple or compound ovary in a flower with only one pistil. Dry fruits may be either dehiscent, or indehiscent. Types of dry, simple fruits, examples of each, include: achene – most seen in aggregate fruits capsule – caryopsis – cypsela – an achene-like fruit derived from the individual florets in a capitulum. Fibrous drupe – follicle – is formed from a single carpel, opens by one suture